Steven J. Seybold
Former Employee
Ecosystem Function and Health Program
1731 Research Park Drive
Davis, CA 95618
United States
Davis, CA 95618
United States
Current Research
Seybold is a research entomologist who specializes in the study of bark and wood-boring beetles. He and his colleagues are characterizing the invasive bark beetle and woodborer fauna of California and other Western U.S. States. The discoveries and descriptions of the life histories and behaviors of these insects are an important first step to understanding their pest status in U.S. forests. Seybold and colleagues also were the first to isolate specific genes from bark beetles that the insects use to detoxify defense chemicals released by trees under attack. The isolation of these genes provides a site for the next generation of bark beetle control tools.
Research Interest
In the future, Seybold plans to explore issues of drought and bark beetle-caused tree mortality, bark beetle and woodborer invasions of California's national forests, and new threats to native oaks and walnuts.
Professional Organizations
- Member, Entomological Society of America, Current
- Member, Entomological Society of Canada, Current
- Member, International Society of Chemical Ecology, Current
Patents
- Seybold, Steve; Dallara, Paul L.; Nelson, Lori; Graves, Andrew D.; Hishinuma, Stacy M; Gries, Regine - 2015 - Methods of Monitoring and Controlling the Walnut Twig Beetle, Pityophthorus Juglandis
Other Publications
- Steven J. Seybold, Christopher J. Fettig. 2021. Managing bark and ambrosia beetles (Coleoptera: Curculionidae: Scolytinae) with semiochemicals
- Barbara Bentz, Pierluigi Pierluigi , Horst Delb, Chris Fettig, Therese Poland, Deepa Pureswaran, Steve Seybold. 2020. Advances in understanding and managing insect pests of forest trees
- Yigen Chen, Brian H. Aukema, Steven J. Seybold. 2020. The effects of weather on the flight of an invasive bark beetle, Pityophthorus juglandis
- Steven Seybold, William Klingeman, Stacy Hishinuma, Tom Coleman, Andrew D. Graves. 2019. Status and impact of walnut twig beetle in urban forest, orchard, and native forest ecosystems
- Tom W. Coleman, Adrian L. Poloni, Yigen Chen, Pham Quang Thu, Qiao Li, Jianghua Sun, Robert J. Rabaglia, Gary Man, Steven J. Seybold. 2019. Hardwood injury and mortality associated with two shot hole borers, Euwallacea spp., in the invaded region of southern California, USA, and the native region of Southeast Asia
- Jesse L Morris, Stuart Cottrell, Christopher J Fettig, R. Justin DeRose, Katherine M Mattor, Vachel A Carter, Jennifer Clear, Jessica Clement, Winslow D Hansen, Jeffrey A Hicke, Philip E Higuera, Alistair WR Seddon, Heikki Seppä, Rosemary L Sherriff, John D Stednick, Steven J Seybold. 2018. Bark beetles as agents of change in social-ecological systems
- Steven J. Seybold, Barbara J. Bentz, Christopher J. Fettig, John E. Lundquist, Robert A. Progar, Nancy E. Gillette. 2018. Management of western North American bark beetles with semiochemicals
- Andrea R. Hefty, Brian H. Aukema, Robert C. Venette, Mark V Coggeshall, James R. McKenna, Steven J. Seybold. 2018. Reproduction and potential range expansion of walnut twig beetle across the Juglandaceae
- Emel Oren, William Klingeman, Romina Gazis, John Moulton, Paris Lambdin, Mark Coggeshall, Jiri Hulcr, Steven J. Seybold, Denita Hadziabdic. 2018. A novel molecular toolkit for rapid detection of the pathogen and primary vector of thousand cankers disease
- Romina Gazis, Laura Poplawski, William Klingeman, Sarah L. Boggess, Robert N. Trigiano, Andrew D. Graves, Steven J. Seybold, Denita Hadziabdic. 2018. Mycobiota associated with insect galleries in walnut with thousand cankers disease reveals a potential natural enemy against Geosmithia morbida
- Aubree M. Kees, Andrea R. Hefty, Robert C. Venette, Steven J. Seybold, Brian H. Aukema. 2017. Flight Capacity of the Walnut Twig Beetle (Coleoptera: Scolytidae) on a Laboratory Flight Mill
- Andrea R. Hefty, Steven J. Seybold, Brian H. Aukema, Robert C. Venette. 2017. Cold Tolerance of Pityophthorus juglandis (Coleoptera: Scolytidae) From Northern California
- Stacy M. Hishinuma, Paul L. Dallara, Mohammad A. Yaghmour, Marcelo M. Zerillo, Corwin M. Parker, Tatiana V. Roubtsova, Tivonne L. Nguyen, Ned A. Tisserat, Richard M. Bostock, Mary L. Flint, Steven J. Seybold. 2016. Wingnut (Juglandaceae) as a new generic host for Pityophthorus juglandis (Coleoptera: Curculionidae) and the thousand cankers disease pathogen, Geosmithia morbida (Ascomycota: Hypocreales)
- Steven J.Seybold, Richard L.Penrose, Andrew Graves. 2016. Invasive bark and ambrosia beetles in California Mediterranean forest ecosystems
- Tom W.Coleman, Steven J.Seybold. 2016. Goldspotted oak borer in California: Invasion history, biology, impact, management, and implications for Mediterranean forests worldwide
- Yigen Chen, Paul L. Dallara, Lori J. Nelson, Tom W. Coleman, Stacy M. Hishinuma, Daniel Carrillo, Steven J. Seybold. 2016. Comparative morphometric and chemical analyses of phenotypes of two invasive ambrosia beetles (Euwallacea spp.) in the United States
- Jesse L. Morris, Stuart Cottrell, Christopher J. Fettig, Winslow D. Hansen, Rosemary L. Sherriff, Vachel A. Carter, Jennifer L. Clear, Jessica Clement, R. Justin DeRose, Jeffrey A. Hicke, Philip E. Higuera, Katherine M. Mattor, Alistair W. R. Seddon, Heikki T. Seppä, John D. Stednick, Steven J. Seybold. 2016. Managing bark beetle impacts on ecosystems and society: priority questions to motivate future research
- Andrea R. Hefty, Mark V. Coggeshall, Brian H. Aukema, Robert C. Venette, Steven J. Seybold. 2016. Reproduction of walnut twig beetle in black walnut and butternut
- Tom W. Coleman, Michael I. Jones, Mark S. Hoddle, Laurel J. Haavik, John C. Moser, Mary L. Flint, Steven J. Seybold. 2015. Pyemotes tritici (Acari: Pyemotidae): a parasitoid of Agrilus auroguttatus and Agrilus coxalis (Coleoptera: Buprestidae) in the southwestern United States of America and southern Mexico
- Paul F. Rugman-Jones, Steven J. Seybold, Andrew D. Graves, Richard Stouthamer. 2015. Phylogeography of the walnut twig beetle, Pityophthorus juglandis, the vector of thousand cankers disease in North American walnut trees
- T.W. Coleman, M.I. Jones, S.L. Smith, R.C. Venette, M.L. Flint, S.J. Seybold. 2015. Goldspotted oak borer, Agrilus auroguttatus
- Laurel J. Haavik, Mary L. Flint, Tom W. Coleman, Robert C. Venette, Steven J. Seybold. 2015. Goldspotted oak borer effects on tree health and colonization patterns at six newly-established sites
- Yigen Chen, Mary L. Flint, Tom W. Coleman, Joseph J. Doccola, Donald M. Grosman, David L. Wood, Steven J. Seybold. 2015. Impact of the goldspotted oak borer, Agrilus auroguttatus, on the health of coast live oak before and after treatment with two systemic insecticides
- Steve Seybold, Tom W. Coleman. 2015. The goldspotted oak borer: revisiting the status of an invasive pest six years after its discovery
- Robert C. Venette, Tom W. Coleman, Steve Seybold. 2015. Assessing the risks posed by goldspotted oak borer to California and beyond
- Yigen Chen, Mary L. Flint, Tom W. Coleman, Joseph J. Doccola, Donald M. Grosman, David L. Wood, Steven J. Seybold. 2015. Effects of the goldspotted oak borer, Agrilus auroguttatus, on the health of coast live oak, Quercus agrifolia, in southern California before and after treatment with two systemic insecticides
- L.J. Haavik, A.D. Graves, T.W. Coleman, F.L. Flint, R.C. Venette, S.J. Seybold. 2014. Suitability of native and ornamental oak species in California for Agrilus auroguttatus
- Laurel J. Haavik, Tom W. Coleman, Mary Louise Flint, Robert C. Venette, Steven J. Seybold. 2014. Densities of Agrilus auroguttatus and other borers in California and Arizona oaks
- Maia M. Beh, Margaret R. Metz, Steven J. Seybold, David M. Rizzo. 2014. The novel interaction between Phytophthora ramorum and wildfire elicits elevated ambrosia beetle landing rates on tanoak, Notholithocarpus densiflorus
- Tom W. Coleman, Michael I. Jones, Beatrice Courtial, Andrew D. Graves, Meghan Woods, Alain Roques, Steven J. Seybold. 2014. Impact of the first recorded outbreak of the Douglas-fir tussock moth, Orgyia pseudotsugata, in southern California and the extent of its distribution in the Pacific Southwest region
- Yigen Chen, Steven J. Seybold. 2014. Crepuscular flight activity of an invasive insect governed by interacting abiotic factors
- Marcelo M. Zerillo, Jorge Ibarra Caballero, Keith Woeste, Andrew D. Graves, Colleen Hartel, Jay W. Pscheidt, Jadelys Tonos, Kirk Broders, Whitney Cranshaw, Steven J. Seybold, Ned Tisserat. 2014. Population structure of Geosmithia morbida, the causal agent of thousand cankers disease of walnut trees in the United States
- Tom W. Coleman, Yigen Chen, Andrew D. Graves, Stacy M. Hishinuma, Nancy E. Grulke, Mary Louise Flint, Steven J. Seybold. 2014. Developing monitoring techniques for the invasive goldspotted oak borer (Coleoptera: Buprestidae) in California
- Michael I. Jones, Tom W. Coleman, Andrew D. Graves, Mary Louise Flint, Steven J. Seybold. 2013. Sanitation options for managing oak wood infested with the invasive goldspotted oak borer (Coleoptera: Buprestidae) in Southern California
- Steven J. Seybold, Paul L. Dallara, Stacy M. Hishinuma, Mary Louise Flint. 2013. Detecting and identifying the walnut twig beetle: Monitoring guidelines for the invasive vector of thousand cankers disease of walnut
- S.J. Seybold, P.L. Dallara, S.M. Hishinuma, M.L. Flint. 2013. Quick guide: Installing, maintaining, and servicing walnut twig beetle pheromone-baited traps.
- Curtis Utley, Tivonne Nguyen, Tatiana Roubtsova, Mark Coggeshall, Tim M. Ford, L.J. Grauke, Andrew D. Graves, Charles A. Leslie, James McKenna, Keith Woeste, Mohammad A. Yaghmour, Steve Seybold, Richard M. Bostock, Ned Tisserat. 2013. Susceptibility of walnut and hickory species to Geosmithia morbida
- L.J. Haavik, T.W. Coleman, M.L. Flint, R.C. Venette, S.J. Seybold. 2013. Agrilus auroguttatus (Coleoptera: Buprestidae) seasonal development within Quercus agrifolia (Fagales: Fagaceae) in southern California
- Y. Chen, S. J. Seybold. 2013. Application of a frequency distribution method for determining instars of the beet armyworm (Lepidoptera: Noctuidae) from widths of cast head capsules
- M.L. Flint, M. I. Jones, T. W. Coleman, S.J. Seybold. 2013. Goldspotted oak borer
- Maia M. Beh, Margaret Metz, Steven J. Seybold, David Rizzo. 2013. The novel interaction between Phytophthora ramorum and wildfire elicits elevated ambrosia beetle landing rates on tanoak
- Yigen Chen, Tom. W. Coleman, Michael. I. Jones, Mary. L. Flint, Steven. J. Seybold. 2013. Foliar nutrients explain goldspotted oak borer, Agrilus auroguttatus, adult feeding preference among four California oak species
- Laurel J. Haavik, Tom W. Coleman, Mary Louise Flint, Robert C. Venette, Steven J. Seybold. 2012. Agrilus auroguttatus exit hole distributions on Quercus agrifolia boles and a sampling method to estimate their density on individual trees
- Paul.L. Dallara, Mary.L. Flint, Steven. J. Seybold. 2012. An analysis of the larval instars of the walnut twig beetle, Pityophthorus juglandis, in northern California black walnut, Juglans hindsii, and a new host record for Hylocurus hirtellus
- Steven J. Seybold, Jennifer A. King, Daren R. Harris, Lori J. Nelson, Shakeeb M. Hamud, Yigen Chen. 2012. Diurnal flight response of the walnut twig beetle, Pityophthorus juglandis, to pheromone-baited traps in two northern California walnut habitats
- Steven J. Seybold, Tom W. Coleman, Paul L. Dallara, Norman L. Dart, Andrew D. Graves, Lee A. Pederson, Sven-Erik Spichiger. 2012. Recent collecting reveals new state records and the extremes in the distribution of the walnut twig beetle, Pityophthorus juglandis Blackman (Coleoptera: Scolytidae), in the United States
- Laurel J. Haavik, Tom W. Coleman, Yigen Chen, Micheal I. Jones, Robert C. Venette, Mary L. Flint, Steven J. Seybold. 2012. First occurrence of the goldspotted oak borer parasitoid, Calosota elongata (Hymenoptera: Eupelmidae), in California
- Steven J. Seybold, Andrew D. Graves, Tom W. Coleman. 2011. Walnut twig beetle: update on the biology and chemical ecology of a vector of an invasive fatal disease of walnut in the western U.S.
- Steven J. Seybold, Tom W. Coleman. 2011. The goldspotted oak borer: an overview of a research program for "California's Emerald Ash Borer"
- Tom W. Coleman, Nancy E. Grulke, Miles Daly, Cesar Godinez, Susan L. Schilling, Philip J. Riggan, Steven J. Seybold. 2011. Coast live oak, Quercus agrifolia, susceptibility and response to goldspotted oak borer, Agrilus auroguttatus, injury in southern California
- S. Hishinuma, T.W. Coleman, M.L. Flint, S.J. Seybold. 2011. Goldspotted oak borer: Field identification guide
- S. Kim, T.C. Harrington, J. C. Lee, S. J. Seybold. 2011. Leptographium tereforme, sp. nov. and other Ophiostomatales isolated from the redhaired pine bark beetle, Hylurgus ligniperda, in California
- T.W. Coleman, Steve Seybold. 2011. Collection history and comparison of the interactions of the goldspotted oak borer, Agrilus auroguttatus Schaeffer (Coleoptera: Buprestidae), with host oaks in southern California and southeastern Arizona
- Jana C. Lee, Jose F. Negron, Sally J. McElwey, Livy Williams, Jeffrey J. Witcosky, John B. Popp, Steven J. Seybold. 2011. Biology of the invasive banded elm bark beetle (Coleoptera: Scolytidae) in the western United States
- N. Tisserat, W. Cranshaw, M. Putnam, J. Pscheidt, C.A. Leslie, M. Murray, J. Hoffman, Y. Barkely, K. Alexander, Steve Seybold. 2011. Thousand cankers disease is widespread on black walnut, Juglans nigra, in the western United States
- J.E. Lundquist, A.E. Camp, M.L. Tyrell, S.J. Seybold, P. Cannon, D.J. Lodge. 2011. Earth, wind, and fire: Abiotic factors and the impacts of global environmental change on forest health
- Roger E. Burnside, Edward H. Holsten, Christopher J. Fettig, James J. Kruse, Mark E. Schultz, Christopher J. Hayes, Andrew D. Graves, Steven J. Seybold. 2011. The northern spruce engraver, Ips perturbatus
- Abigail J. Walter, Stephen A. Kells, Robert C. Venette, Steven J. Seybold. 2010. Boring in response to bark and phloem extracts from North American trees does not explain host acceptance behavior of Orthotomicus erosus (Coleoptera: Scolytidae)
- Tom W. Coleman, Steven J. Seybold. 2010. GSOB ≠ SOD. Tree mortality from the goldspotted oak borer in oak woodlands of southern California.
- Jana C. Lee, Shakeeb M. Hamud, Jose F. Negron, Jeffrey J. Witcosky, Steven J. Seybold. 2010. Semiochemical-mediated flight strategies of two invasive elm bark beetles: A potential factor in competitive displacement
- Barbara J. Bentz, Jacques Regniere, Christopher J. Fettig, E. Matthew Hansen, Jane L. Hayes, Jeffrey A. Hicke, Rick G. Kelsey, Jose F. Negron, Steven J. Seybold. 2010. Climate change and bark beetles of the western United States and Canada: Direct and indirect effects
- K. J. Dodds, D. W. Gilmore, S. J. Seybold. 2010. Assessing the threat posed by indigenous exotics: A case study of two North American bark beetle species
- A. J. Walter, R. C. Venette, S. A. Kells, S. J. Seybold. 2010. Walking response of the Mediterranean pine engraver, Orthotomicus erosus, to novel plant odors host in a laboratory olfactometer
- T.W. Coleman, S.J. Seybold. 2010. Verification of a useful character for separating the sexes of the goldspotted oak borer, Agrilus coxalis auroguttatus (Coleoptera: Buprestidae)
- V.R. Lewis, S.J. Seybold. 2010. Wood-boring beetles in homes
- Yilmaz Balci, William L. MacDonald, Kurt Gottschalk, Elisa Becker, Grace Sumampong, Simon F. Shamoun, Aniko Varga, Saad Masri, Delano James, Karen Bailey, Susan Boyetchko, Maia M. Beh, Kerri Frangioso, Akiko Oguchi, Guillaume J. Bilodeau, Richard C. Hamelin, Francoise Pelletier, C. Andre Levesque, Jennifer Britt, Everett Hansen, Monica Bueno, Janelle Deshais, Leonel Arguello, Gary Chastagner, Annie DeBauw, Kathy Riley, Marianne Elliott, Ellen Crocker, Matteo Garbelotto, John Stuart, Steven Steinberg, Clare R. Elliott, Virginia McDonald, Katie Coats, Niklaus J. Grunwald, Elizabeth Fichtner, Tedmund J. Swiecki, Elizabeth A. Bernhardt, Allison Wickland, David Rizzo, Ross Meentemeyer, S.E. Haas, M. Metz, Timothy L. Widmer, Gary J. Samuels, Paul W. Tooley, Marsha E. Browning, Harvey Timeus, Wendy Sutton, Paul Reeser, Alan Kanaskie, Steven J. Seybold, Tom W. Coleman, Mary Louise Flint, T. Henkel, A. Breitenstein, S. Werres, M. Riedel, S. Julich, M. Kielpinski, R. Möller, W. Fritsche, S. Wagner, Michael Thompson, Ellen Michaels Goheen, Jon Laine, Nathan Rank, Hall Cushman, Jennifer L. Parke, Jay Pscheidt, Richard Regan, Jan Hedberg, Steven W. Oak, Jaesoon Hwang, Steven N. Jeffers. 2010. Posters
- Jana Lee, Ingrid Aguayo, Ray Aslin, Gail Durham, Shakeeb Hamud, Bruce Moltzan, Steve Munson, Jose Negron, Travis Peterson, Iral Ragenovich, Jeff Witcosky, Steven Seybold. 2009. Trend of two invasives: the banded and European elm bark beetles
- Janna C. Lee, Ingrid Aguayo, Ray Aslin, Gail Durham, Shakeeb M. Hamud, Beruce D. Moltzan, A. Steve Munson, Jose F. Negron, Travis Peterson, Iral R. Ragenovich, Jeffrey J. Witcosky, Steven J. Seybold. 2009. Co-occurrence of the invasive banded and European elm bark beetles (Coleoptera: Scolytidae) in North America
- N.E. Grulke, R.A. Minnich, T.D. Paine, S.J. Seybold, D.J. Chavez, M.E. Fenn, P.J. Riggan, A. Dunn. 2009. Air pollution increases forest susceptibility to wildfires: A case study in the San Bernardino Mountains in southern California
- S.J. Seybold, M. Downing. 2009. What risk do invasive bark beetles and woodborers pose to forests of the western U.S.?: A case study of the Mediterranean pine engraver, Orthotomicus erosus
- Barbara Bentz, Jesse Logan, Jim MacMahon, Craig D. Allen, Matt Ayres, Ed Berg, Allan Carroll, Matt Hansen, Jeff Hicke, Linda Joyce, Wallace Macfarlane, Steve Munson, Jose Negron, Tim Paine, Jim Powell, Ken Raffa, Jacques Regniere, Mary Reid, Bill Romme, Steven J. Seybold, Diana Six, Diana Tomback, Jim Vandygriff, Tom Veblen, Mike White, Jeff Witcosky, David Wood. 2009. Bark beetle outbreaks in western North America: Causes and consequences
- J.C. Lee, S.J. Seybold. 2009. Host acceptance and larval competition between the invasive banded and European elm bark beetles (Coleoptera: Scolytidae): Potential mechanisms for competitive displacement
- C.J. Fettig, S.R. McKelvey, R.R. Borys, C.P Dabney, S.M. Hamud, L.J. Nelson, S.J. Seybold. 2009. Efficacy of verbenone for protecting ponderosa pine stands from western pine beetle (Coleoptera: Curculionidae, Scolytinae) attack in California
- Tom W. Coleman, Steven J. Seybold. 2009. Striking gold in southern California: discovery of the goldspotted oak borer and its central role in oak mortality
- K.J.K. Gandhi, D.W. Gilmore, R.A. Haack, S.A. Katovich, S.J. Krauth, W.J. Mattson, J.C. Zasada, S.J. Seybold. 2009. Application of semiochemicals to assess the biodiversity of subcortical insects following an ecosystem disturbance in a sub-boreal forest
- Kier D. Klepzig, Therese M. Poland, Nancy E. Gillette, Robert A. Haack, Melody A. Keena, Daniel R. Miller, Michael E. Montgomery, Steven J. Seybold, Patrick C. Tobin. 2009. Forest Service R&D — Invasive Insects: Visions for the Future
- Jose F. Negron, Barbara J. Bentz, Christopher J. Fettig, Nancy Gillette, E. Matthew Hansen, Jane L. Hayes, Rick G. Kelsey, John E. Lundquist, Ann M. Lynch, Robert A. Progar, Steven J. Seybold. 2008. US Forest Service bark beetle research in the western United States: Looking toward the future
- Christopher J. Fettig, Joel D. McMillin, John. A. Anhold, Shakeeb M. Hamud, Steven J. Seybold, Robert R. Borys. 2008. The effects of hazardous fuel reduction treatments in the wildland urban interface on the activity of bark beetles infesting ponderosa pine
- Jana C. Lee, Shakeeb Hamud, Jose Negron, Jeff Witcosky, A. Steve Munson, Steven J. Seybold. 2008. The exotic banded elm bark beetle and the smaller European beetle: abundance host attraction
- Kamal J.K. Gandhi, Daniel W. Gilmore, Steven A. Katovich, William J. Mattson, John C. Zasada, Steven J. Seybold. 2008. Catastrophic windstorm and fuel-reduction treatments alter ground beetle (Coleoptera: Carabidae) assemblages in a North American sub-boreal forest
- Tom W. Coleman, Steven Seybold. 2008. Previously unrecorded damage to oak, Quercus spp., in southern California by the goldspotted oak borer, Agrilus coxalis Waterhouse (Coleoptera: Buprestidae)
- William Shepherd, Dezene Huber, Steven Seybold, Christopher Fettig. 2008. Antennal responses of the western pine beetle, Dendroctonus brevicomis (Coleoptera: Curculionidae), to stem volatiles of its primary host, Pinus ponderosa, and nine sympatric nonhost angiosperms and conifers.
- Dezene P. W. Huber, Melissa Erickson, Christian Leutenegger, Joerg Bohlmann, Steven J. Seybold. 2007. Expression patterns of bark beetle cytochromes P450 during host colonization: Likely physiological functions and potential targets for pest management
- Christopher J. Fettig, Joel D. McMillin, John A. Anhold, Shakeeb M. Hamud, Robert R. Borys, Steven J. Seybold. 2007. The effect of mechanical fuel reduction treatments in the wildland-urban interface on the amount and distribution of bark beetle-caused tree mortality
- Deguang Liu, Michael J. Bohne, Jana C. Lee, Mary Louise Flint, Richard L. Penrose, Steven J. Seybold. 2007. New introduction in California: the redhaired pine bark beetle, Hylurgus ligniperda Fabricius
- Jose F. Negron, Jeffrey J. Witcosky, Robert J. Cain, James R. LaBonte, Donald A. II.Duerr, Sally J. McElwey, Jana C. Lee, Steven J. Seybold. 2005. The banded elm bark beetle: A new threat to elms in North America
- Jana C. Lee, Sheri L. Smith, Steven J. Seybold. 2005. The Mediterranean pine engraver
- Robert J. Bartelt, John F. Kyhl, Angie K. Ambourn, Jennifer Juzwik, Steven J. Seybold. 2004. Male-produced aggregation pheromone of Carpophilus sayi, a nitidulid vector of oak wilt disease, and pheromonal comparison with Carpophilus lugubris
- John F. Kyhl, Robert J. Bartelt, Allard Cosse, Jennifer Juzwik, Steven J. Seybold. 2002. Semiochemical-mediated flight responses of sap beetle vectors of oak wilt, Ceratocystis fagacearum
- Edward H. Holsten, Roger E. Burnside, Steven J. Seybold. 2000. Attractant semiochemicals of the engraver beetle, Ips perturbatus, in south-central and interior Alaska.
- Daniel R. Miller, Kenneth E. Gibson, K.F. Raffa, S.J. Seybold, S.A. Teale, D.L. Wood. 1997. Geographic variation in response of pine engraver, Ips pini, and associated species to pheromone, lanierone
- Steven J. Seybold. 1992. The Status of Scolytid Genetics-A Reductionist's View
- David L. Wood, Andrew J. Storer, Thomas R. Gordon, Steven J. Seybold, Marion Page. 1992. Biosystematics of Ips mexicanus and Ips plastographus (Coleoptera: Scolytidae) and Their Fungal Symbionts