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    The purpose of this study was to determine the most important nonanthropogenic factors affecting understory (herbs, shrubs and low-growing vines) diversity in forested landscapes of southern Indiana. Fourteen environmental variables were measured for 46 sites. Multiple regression analysis showed significant positive correlation between understory diversity and tree seedling diversity and soil subgroup. Slope aspect, soil rooting depth and number of standing dead trees were significantly negatively correlated with understory diversity. Mesic sites were more diverse in common understory species than xeric sites but had lower total cover and different species.

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    Huebner, Cynthia D.; Randolph, J.C.; Parker, G.R. 1995. Environmental factors affecting understory diversity in second-growth deciduous forests. American Midland Naturalist. 134: 155-165.

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