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    The higher members of the tribe Attini characteristically lay persistent and extensive odor trails especially in many neotropical areas. Thus, in Acromyrmex and Atta, long columns of foraging workers are frequently present on the odor trails but in the less specialized attine genera, workers may forage either in files or singly. Weber (1958) has indicated that the workers in monomorphic genera forage singly, a behavioral characteristic which he offers as evidence for the primitive position of these attine genera.

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    Blum, Murray S.; Moser, John C.; Cordero, A.D. 1964. Chemical releases of social behavior, II. source and specificity of the odor trail substances in four attine genera. (Hymenoptera: Formicidae). Psyche, Vol. 71(1): 1-7

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