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Widespread increase of tree mortality rates in the Western United States


Phillip J. van Mantgem
Nathan L. Stephenson
Lori D. Daniels
Jerry F. Franklin
Peter Z. Fule
Mark E. Harmon
Andrew J. Larson
Jeremy M. Smith
Alan H. Taylor
Thomas T. Veblen



Publication type:

Scientific Journal (JRNL)

Primary Station(s):

Pacific Northwest Research Station



DOI: 10.1126/science.1165000


Persistent changes in tree mortality rates can alter forest structure, composition, and ecosystem services such as carbon sequestration. Our analyses of longitudinal data from unmanaged old forests in the Western United States showed that background (noncatastrophic) mortality rates have increased rapidly in recent decades, with doubling periods ranging from 17 to 29 years among regions. Increases were also pervasive across elevations, tree sizes, dominant genera, and past fire histories. Forest density and basal area declined slightly, which suggests that increasing mortality was not caused by endogenous increases in competition. Because mortality increased in small trees, the overall increase in mortality rates cannot be attributed solely to aging of large trees. Regional warming and consequent increases in water deficits are likely contributors to the increases in tree mortality rates.


van Mantgem, Phillip J.; Stephenson, Nathan L.; Byrne, John C.; Daniels, Lori D.; Franklin, Jerry F; Fule, Peter Z; Harmon, Mark E.; Larson, Andrew J.; Smith, Jeremy M.; Taylor, Alan H.; Veblen, Thomas T. 2009. Widespread increase of tree mortality rates in the Western United States. Science. 323:521-524.

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