Index of Species Information
SPECIES: Bromus ciliatus
Introductory
SPECIES: Bromus ciliatus
AUTHORSHIP AND CITATION :
Esser, Lora L. 1994. Bromus ciliatus. In: Fire Effects Information System, [Online].
U.S. Department of Agriculture, Forest Service,
Rocky Mountain Research Station, Fire Sciences Laboratory (Producer).
Available: https://www.fs.usda.gov
/database/feis/plants/graminoid/brocil/all.html [].
ABBREVIATION :
BROCIL
SYNONYMS :
Bromopsis ciliatus (L.) Holub [59]
SCS PLANT CODE :
BRCI2
COMMON NAMES :
fringed brome
fringed bromegrass
TAXONOMY :
The currently accepted scientific name of fringed brome is Bromus
ciliatus L. [37,59]. It is a member of the Poaceae family. There
are two recognized forms [66]:
B. c. f. ciliatus
B. c. f. intonus (Fern.) Seymour
Some authorities classify the tetraploid race of this plant as a
distinct species, B. richardsonii Link [27,32].
LIFE FORM :
Graminoid
FEDERAL LEGAL STATUS :
No special status
OTHER STATUS :
NO-ENTRY
DISTRIBUTION AND OCCURRENCE
SPECIES: Bromus ciliatus
GENERAL DISTRIBUTION :
Fringed brome occurs from Alaska south to southern California and east
throughout the western states, the Great Plains, and discontinuously
through the midwestern, northeastern, and Atlantic coastal states
[24,29,46,59]. It occurs throughout Canada, excluding the Northwest
Territories and Prince Edward Island [23,24,37]. Fringed brome also
occurs in Mexico [29,59].
ECOSYSTEMS :
FRES10 White - red - jack pine
FRES11 Spruce - fir
FRES17 Elm - ash - cottonwood
FRES18 Maple - beech - birch
FRES19 Aspen - birch
FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES23 Fir - spruce
FRES26 Lodgepole pine
FRES28 Western hardwoods
FRES29 Sagebrush
FRES34 Chaparral - mountain shrub
FRES35 Pinyon - juniper
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES38 Plains grasslands
FRES39 Prairie
STATES :
AK CA CO ID IL IN IA KS ME MA
MI MN MO MT NE NH NJ NM NV NY
NC ND OK OR PA RI SD TN TX UT
VT VA WA WI WV WY AB BC MB NB
NF NS ON PQ SK YT MEXICO
BLM PHYSIOGRAPHIC REGIONS :
1 Northern Pacific Border
2 Cascade Mountains
3 Southern Pacific Border
4 Sierra Mountains
5 Columbia Plateau
6 Upper Basin and Range
7 Lower Basin and Range
8 Northern Rocky Mountains
9 Middle Rocky Mountains
10 Wyoming Basin
11 Southern Rocky Mountains
12 Colorado Plateau
13 Rocky Mountain Piedmont
14 Great Plains
15 Black Hills Uplift
16 Upper Missouri Basin and Broken Lands
KUCHLER PLANT ASSOCIATIONS :
K005 Mixed conifer forest
K008 Lodgepole pine - subalpine forest
K010 Ponderosa shrub forest
K011 Western ponderosa forest
K012 Douglas-fir forest
K015 Western spruce - fir forest
K016 Eastern ponderosa forest
K017 Black Hills pine forest
K018 Pine - Douglas-fir forest
K019 Arizona pine forest
K020 Spruce - fir - Douglas-fir forest
K021 Southwestern spruce - fir forest
K022 Great Basin pine forest
K023 Juniper - pinyon woodland
K026 Oregon oakwoods
K031 Oak - juniper woodlands
K033 Chaparral
K037 Mountain-mahogany - oak scrub
K050 Fescue - wheatgrass
K055 Sagebrush steppe
K066 Wheatgrass - needlegrass
K074 Bluestem prairie
K093 Great Lakes spruce - fir forest
K094 Conifer bog
K095 Great Lakes pine forest
K101 Elm - ash forest
K107 Northern hardwoods - fir forest
SAF COVER TYPES :
1 Jack pine
5 Balsam fir
12 Black spruce
13 Black spruce - tamarack
16 Aspen
18 Paper birch
21 Eastern white pine
37 Northern white-cedar
38 Tamarack
63 Cottonwood
107 White spruce
108 Red maple
201 White spruce
202 White spruce - paper birch
203 Balsam poplar
204 Black spruce
206 Engelmann spruce - subalpine fir
209 Bristlecone pine
210 Interior Douglas-fir
211 White fir
216 Blue spruce
217 Aspen
218 Lodgepole pine
219 Limber pine
220 Rocky Mountain juniper
233 Oregon white oak
235 Cottonwood - willow
237 Interior ponderosa pine
239 Pinyon - juniper
243 Sierra Nevada mixed conifer
244 Pacific ponderosa pine - Douglas-fir
245 Pacific ponderosa pine
251 White spruce - aspen
252 Paper birch
253 Black spruce - white spruce
254 Black spruce - paper birch
SRM (RANGELAND) COVER TYPES :
NO-ENTRY
HABITAT TYPES AND PLANT COMMUNITIES :
Fringed brome occurs in a wide variety of habitat types including
pinyon-juniper (Pinus spp.-Juniperus spp.), quaking aspen (Populus
tremuloides), fir-spruce (Abies spp.-Picea spp.), subalpine fir (Abies
lasiocarpa), ponderosa pine (Pinus ponderosa), lodgepole pine (P.
contorta), Douglas-fir (Pseudotsuga menziesii), maple (Acer spp.),
sagebrush, grassland, and riparian [1,6,11,40,59]. It is an indicator
of aspen and riparian community types in the Intermountain region
[11,43,45].
Fringed brome occurs in coniferous forest, in both climax and seral
communities. A Douglas-fir/fringed brome habitat type has been
described for northern New Mexico and northern Arizona [1,20,38].
Fringed brome is a member of the single-leaf pinyon-Utah juniper (Pinus
monophylla-J. osteosperma) association of northern Arizona [13].
Fringed brome is a dominant understory species in subalpine fir,
lodgepole pine, white fir (Abies concolor), and blue spruce (Picea
pungens) habitat types of Arizona and New Mexico [1,2,20,43]. It is a
common understory species in subalpine fir and Engelmann spruce (P.
engelmannii) habitat types of Idaho and western Wyoming [11].
Fringed brome also occurs in a number of other communities. It is found
in seral aspen community types, including the aspen/fringed brome
community typed described for Utah [48] and aspen-dominated community
types in Wyoming, Colorado, and New Mexico [45]. Fringed brome is a
member of the sedge-goldenrod (Carex spp.-Solidago spp.) wet prairie
community in Michigan [33]. It also occurs in peatlands of
north-central Minnesota in minerotrophic fens [60].
In Canada, fringed brome is an understory species in a 100-year-old
black spruce (Picea mariana) forest in southeastern Manitoba [14]. It
also occurs in stagnant plantations of prethicket white spruce (P.
glauca) on oldfield sites in Ontario [56]. Fringed brome is a member
of the western snowberry (Symphoricarpos occidentalis) shrub community
in Alberta [5].
The following publications list fringed brome as a community dominant:
A classification of forest habitat types of the northern portion of the
Cibola National Forest, New Mexico [1]
Forest habitat types in the Apache, Gila, and part of the Cibola
National Forests, Arizona and New Mexico [20]
Forest and woodland habitat types (plant associations) of northern New
Mexico and northern Arizona [38]
A classification of spruce-fir and mixed conifer habitat types of
Arizona and New Mexico [43]
Aspen community types of the Pike and San Isabel National Forests in
south-central Colorado [48]
Species not previously mentioned but commonly associated with fringed
brome in New Mexico include Arizona white oak (Quercus arizonica),
Gambel oak (Q. gambelii), silverleaf oak (Q. hypoleucoides), alligator
juniper (Juniperus deppeana), Arizona walnut (Juglans major), Arizona
madrone (Arbutus arizonica), sacahuista (Nolina microcarpa), skunkbush
sumac (Rhus trilobata), mountain snowberry (Symphoricarpos oreophilus),
dwarf bilberry (Vaccinium myrtillus), desert ceanothus (Ceanothus
greggii), willow (Salix spp.), Oregon-grape (Mahonia repens), western
yarrow (Achillea millefolium), bush oceanspray (Holodiscus dumosus),
Fendler meadowrue (Thalictrum fendleri), New Mexico locust (Robinia
neomexicana), green ephedra (Ephedra viridis), broom snakeweed
(Gutierrezia sarothrae), Ross sedge (Carex rossii), Thurber fescue
(Fescue thurberi), Arizona fescue (F. arizonica), and Wolf currant
(Ribes wolfii) [1,13,43].
Species not previously mentioned but commonly associated with fringed
brome in Utah and Wyoming include narrow-leaved cottonwood (Populus
angustifolium), black sage (Artemisia nova), serviceberry (Amelanchier
spp.), sticky geranium (Geranium viscosissimum), lupine (Lupinus spp.),
woodland strawberry (Fragaria vesca), fireweed (Epilobium
angustifolium), shrubby cinquefoil (Potentilla fruticosa), northern
bedstraw (Galium boreale), Fendler bluegrass (Poa fendleriana), and
slender wheatgrass (Elymus trachycaulus) [3,6,12].
MANAGEMENT CONSIDERATIONS
SPECIES: Bromus ciliatus
IMPORTANCE TO LIVESTOCK AND WILDLIFE :
Fringed brome is a good source of forage on western forest ranges. In
Montana and Wyoming, fringed brome is browsed by livestock and is
considered one of the best range grasses in Wyoming [10,40]. In Arizona
and New Mexico, fringed brome is an important forage species for
livestock, deer, and elk throughout the summer [31,38]. Seeds are eaten
by small mammals, turkeys and other birds [25,38]. The aspen/fringed
brome habitat type of central Colorado is ideal for pocket gophers [48].
PALATABILITY :
Fringed brome is highly palatable to deer, elk, and all classes of
livestock [10,31,59].
NUTRITIONAL VALUE :
Energy rating is fair and protein content is poor for fringed brome
[10].
COVER VALUE :
The white fir-Arizona walnut habitat type of Arizona and New Mexico, in
which fringed brome occurs, provides excellent cover for rodents,
turkeys, and quail. The Douglas-fir/fringed brome habitat type provides
nesting cover for band-tailed pigeons [20].
VALUE FOR REHABILITATION OF DISTURBED SITES :
Erosion control and short-term revegetation potential are rated medium,
and long-term revegetation potential is rated high for fringed brome
[10].
OTHER USES AND VALUES :
NO-ENTRY
OTHER MANAGEMENT CONSIDERATIONS :
The effects of clearcutting on wildlife habitat were studied in a moist
subalpine forest in central Colorado. Fringed brome cover increased 1
year after harvest, and then fluctuated in postharvest years 2 to 5.
Percent understory cover of fringed brome before and after clearcutting
were [15]:
before logging years after logging (1978-1982)
(1976) 1 2 3 4 5
2.9 3.3 1.0 1.6 2.5 1.5
Fringed brome is highly palatable, so it is often grazed too closely.
Humphrey [31] recommended that about 33 percent of the seedstalks be
left ungrazed each year and a grazing deferment every third year be
instituted for continuous availability of fringed brome.
BOTANICAL AND ECOLOGICAL CHARACTERISTICS
SPECIES: Bromus ciliatus
GENERAL BOTANICAL CHARACTERISTICS :
Fringed brome is a nonrhizomatous, native perennial that is generally
tufted [21,24,27]. Culms are slender, usually 1.7 to 4 feet (0.5-1.2 m)
tall, but up to 5.2 feet (1.6 m) tall in the Great Plains [24,59]. The
blades are flat, 0.12 to 0.6 inch (3-15 mm) wide and 6 to 10 inches
(15-25 cm) long [25,59]. The panicle is narrowly elongate, 2.8 to 7.2
inches (7-18 cm) long with branches ascending to drooping [24,46].
Fringed brome has a well-developed root system [25].
RAUNKIAER LIFE FORM :
Hemicryptophyte
REGENERATION PROCESSES :
Fringed brome reproduces exclusively from seed [25]. Seeds are
nondormant and can show high germination rates. Tests were conducted by
Hoffman [30] on herbaceous plants common in aspen understories of
Colorado. Fringed brome exhibited a wide range of germination capacity.
Tests were conducted with light and dark regimes, with or without
stratification, and with a variety of thermoperiods. In 9 tests out of
14, fringed brome had 100 percent germination. The lowest germination
rate recorded was 60 percent in a test in which light followed
stratification [30].
Fringed brome is wind pollinated [25].
SITE CHARACTERISTICS :
Fringed brome occurs in a variety of habitats including woodlands,
forest openings, thickets, grasslands, shrublands, prairies, meadows,
marshes, bogs, fens, and stream and lake margins [24,31,40,48,51,]. It
is commonly found in moist places such as wet meadows, benches, and
along streams [46,58]. Fringed brome also occurs on moist to seasonally
dry, open or densely shaded habitats in valleys and montane zones
[20,36].
Fringed brome grows best on moist to semiwet soils, but is tolerant of
poorly drained and subirrigated conditions [13,40,54]. It grows best on
loam, silty loam, and sand, but occurs on stony or bouldery substrates
as well [13,15,39,59]. Soil pH ranges from 4.8 to 7.9 in Yellowstone
National Park [40].
Elevations for fringed brome for several states and provinces are as
follows:
Arizona 6,000-11,000 feet (1,800-3,300 m) [7,43]
California 3,630-10,560 feet (1,100-3,200 m) [27]
Colorado 6,000-11,000 feet (2,700-3,350 m) [15,26]
Michigan 600-750 feet (180-225 m) [33]
New Mexico 7,500-12,000 feet (2,285-3,600 m) [17,43]
New York 1,486-2,800 feet (445-840 m) [35]
Utah 5,015-11,580 feet (1,520-3,510 m) [59]
Wyoming 7,900 feet (2,400 m) [6]
Ontario 1,551 feet (470 m) [54]
Yukon Territory 2,440 feet (740 m) [52]
SUCCESSIONAL STATUS :
Facultative Seral Species
Fringed brome occurs in seral and climax communities. It is shade
tolerant, but also grows in some open habitats [35,36,40]. It is a
facultative wetland species in Montana [10]. The aspen/fringed brome
community type is successional to coniferous climax types in subalpine
forests of Utah [47]. After major disturbances on Douglas-fir/fringed
brome habitat type sites of New Mexico and Arizona, fringed brome
quickly dominates the understory under aspen [1]. In heavily shaded
microsites in white fir/screwleaf muhly (Muhlenbergia virescens) habitat
types, fringed brome may become the dominant understory species [20].
In the Rocky Mountains of New Mexico, fringed brome is found in
old-growth, intermediate-aged, and young-growth forests [17].
SEASONAL DEVELOPMENT :
Fringed brome flowers from July to August in California, the Upper Great
Plains, and New York [24,35,46]. It flowers in August in Tennessee,
North Carolina, and West Virginia [49]. In Utah, fringed brome flowers
in August and September [3]. It flowers from July to October in Arizona
[32].
FIRE ECOLOGY
SPECIES: Bromus ciliatus
FIRE ECOLOGY OR ADAPTATIONS :
Fringed brome has low tolerance to fires of moderate and high intensity
[6]. The seeds of most plants can survive grass fires [67], so it is
likely that seeds of fringed brome in grasslands survives fire. Seed
survival is increased if the seeds are covered by soil [67].
FIRE REGIMES :
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find Fire Regimes".
POSTFIRE REGENERATION STRATEGY :
Tussock graminoid
Secondary colonizer - off-site seed
FIRE EFFECTS
SPECIES: Bromus ciliatus
IMMEDIATE FIRE EFFECT ON PLANT :
Fringed brome is probably top-killed by most fires, as are most grasses.
In northwestern Wyoming, fringed brome was "harmed" by fires of moderate
and high intensity. Moderate intensity fires killed most surface
vegetation but did not remove all litter and duff and killed less than
90 percent of mature aspens. After 3 years, fringed brome did not show
"appreciable recovery" from the fires [6].
DISCUSSION AND QUALIFICATION OF FIRE EFFECT :
NO-ENTRY
PLANT RESPONSE TO FIRE :
Frequency and cover of fringed brome increased the third year following
a spring prescribed fire in a western snowberry shrub community in
central Alberta. The increase was greater on the unburned plot than the
burned plot [5].
At Breakneck Ridge, Wyoming, a prescribed fire was conducted on August
29, 1974. Fringed brome and slender wheatgrass were the two most
abundant grasses on the site. Fire intensities were classed as follows:
light (<20% of vegetation consumed and very few overstory trees killed),
moderate (21-80% of fine fuels and lesser vegetation consumed by fire
and up to 90% of the mature trees killed), and high (81-100% of fine
fuels consumed and over 90% of overstory trees killed) [6,64]. On the
moderate-intensity burn site, grass cover decreased from 15 to 10
percent by postfire year 3; on the high-intensity burn site, a 66
percent drop occurred by postfire year 3. Fringed brome was affected
most by the fire. By the end of postfire year 3, fringed brome had not
shown an appreciable recovery on moderate or high intensity burn sites
[64,65]. Production (air-dry kg/ha) of fringed brome before burning and
in postfire year 3 on plots burned at three levels of intensity were as
follows [6,64]:
before burning light intensity moderate intensity high intensity
217 267 181 87
In southern Ontario, a low-intensity prescribed surface fire in an aspen
(Populus spp.) woodland was conducted on two plots on May 8, 1972 and
two more were burned on April 24, 1973. All plots were sampled in
August of 1973. Fringed brome only occurred on the control sites, at
2.5 percent frequency [54].
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE :
The Research Project Summary Vegetation recovery following a mixed-severity
fire in aspen groves of western Wyoming provides information on prescribed
fire use and postfire response of plant community species, including fringed
brome, that was not available when this species review was originally written.
FIRE MANAGEMENT CONSIDERATIONS :
Grass fires in Douglas-fir/Arizona fescue habitats of which fringed
brome is a member reduced the density of conifer seedlings and
maintained grass cover, although specific effects on fringed brome were
not described [20].
Aspen/fringed brome stands in subalpine forests of Utah have a moderate
probability of being successfully prescribed burned, but only if
livestock grazing is deferred for at least one season before burning.
Postfire communities "quickly resemble prefire ones" [48].
REFERENCES
SPECIES: Bromus ciliatus
REFERENCES :
1. Alexander, Billy G., Jr.; Fitzhugh, E. Lee; Ronco, Frank, Jr.; Ludwig,
John A. 1987. A classification of forest habitat types of the northern
portion of the Cibola National Forest, New Mexico. Gen. Tech. Rep.
RM-143. Fort Collins, CO: U.S. Department of Agriculture, Forest
Service, Rocky Mountain Forest and Range Experiment Station. 35 p.
[4207]
2. Alexander, Robert R.; Ronco, Frank, Jr. 1987. Classification of the
forest vegetation on the National Forests of Arizona and New Mexico.
Res. Note RM-469. Fort Collins, CO: U.S. Department of Agriculture,
Forest Service, Rocky Mountain Forest and Range Experiment Station. 10
p. [3515]
3. Allman, Verl Phillips. 1953. A preliminary study of the vegetation in an
exclosoure in the chaparral of the Wasatch Mountains, Utah. Utah Academy
Proceedings. 30: 63-78. [9096]
4. Allred, Kelly W. 1993. Bromus, section Pnigma, in New Mexico, with a key
to the bromegrasses of the state. Phytologia. 74(4): 317-318. [23031]
5. Anderson, Murray L.; Bailey, Arthur W. 1979. Effect of fire on a
Symphoricarpos occidentalis shrub community in central Alberta. Canadian
Journal of Botany. 57: 2820-2823. [2867]
6. Bartos, D. L.; Mueggler, W. F. 1981. Early succession in aspen
communities following fire in western Wyoming. Journal of Range
Management. 34(4): 315-318. [5100]
7. Bassett, R.; Larson, M.; Moir, W. 1987. Forest and woodland habitat
types (plant associations) of Arizona south of the Mogollon Rim and
southwestern New Mexico. 2nd Edition. Albuquerque, NM: U.S. Department
of Agriculture, Forest Service, Southwestern Region. [Pages unknown].
[20308]
8. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals,
reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's
associations for the eleven western states. Tech. Note 301. Denver, CO:
U.S. Department of the Interior, Bureau of Land Management. 169 p.
[434]
9. Betters, David R. 1983. Overstory-understory relationships: aspen
forests. In: Bartlett, E.T.; Betters, David R., eds.
Overstory-understory relationships in Western forests. Western Regional
Research Publication No. 1. Fort Collins, CO: Colorado State University
Experiment Station: 5-8. [3309]
10. Boggs, Keith; Hansen, Paul; Pfister, Robert; Joy, John. 1990.
Classification and management of riparian and wetland sites in
northwestern Montana. Missoula, MT: University of Montana, School of
Forestry, Montana Forest and Conservation Experiment Station, Montana
Riparian Association. 217 p. Draft Version 1. [8447]
11. Bradley, Anne F.; Fischer, William C.; Noste, Nonan V. 1992. Fire
ecology of the forest habitat types of eastern Idaho and western
Wyoming. Gen. Tech. Rep. INT-290. Ogden, UT: U.S. Department of
Agriculture, Forest Service, Intermountain Research Station. 92 p.
[19558]
12. Buchanan, Hayle. 1960. The plant ecology of Bryce Canyon National Park.
Salt Lake City, UT: University of Utah. 136 p. Thesis. [3364]
13. Butterwick, Mary; Parfitt, Bruce D.; Hillyard, Deborah. 1992. Vascular
plants of the northern Hualapai Mountains, Arizona. Journal of the
Arizona-Nevada Academy of Science. 24-25: 31-49. [18327]
14. Chrosciewicz, Z. 1976. Burning for black spruce regeneration on a
lowland cutover site in southeastern Manitoba. Canadian Journal of
Forest Research. 6(2): 179-186. [7280]
15. Crouch, Glenn L. 1985. Effects of clearcutting a subalpine forest in
central Colorado on wildlife habitat. Res. Pap. RM-258. Fort Collins,
CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain
Forest and Range Experiment Station. 12 p. [8225]
16. Crouch, Glenn L. 1986. Effects of thinning pole-sized lodgepole pine on
understory vegetation and large herbivore activity in central Colorado.
Res. Pap. RM-268. Fort Collins, CO: U.S. Department of Agriculture,
Forest Service, Rocky Mountain Forest and Range Experiment Station. 10
p. [8353]
17. Dye, A. J.; Moir, W. H. 1977. Spruce-fir forest at its southern
distribution in the Rocky Mountains, New Mexico. American Midland
Naturalist. 97(1): 133-146. [7476]
18. Eyre, F. H., ed. 1980. Forest cover types of the United States and
Canada. Washington, DC: Society of American Foresters. 148 p. [905]
19. Fechner, Gilbert H. 1985. Silvical characteristics of blue spruce. Gen.
Tech. Rep. RM-117. Fort Collins, CO: U.S. Department of Agriculture,
Forest Service, Rocky Mountain Forest and Range Experiment Station. 19
p. [7478]
20. Fitzhugh, E. Lee; Moir, William H.; Ludwig, John A.; Ronco, Frank, Jr.
1987. Forest habitat types in the Apache, Gila, and part of the Cibola
National Forests, Arizona and New Mexico. Gen. Tech. Rep. RM-145. Fort
Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky
Mountain Forest and Range Experiment Station. 116 p. [4206]
21. Fulbright, Timothy E.; Redente, Edward F.; Hargis, Norman E. 1982.
Growing Colorado plants from seed: a state of the art: Volume II:
Grasses and grasslike plants. FWS/OBS-82/29. Washington, DC: U.S.
Department of the Interior, Fish and Wildlife Service. 113 p. [3709]
22. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others].
1977. Vegetation and environmental features of forest and range
ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of
Agriculture, Forest Service. 68 p. [998]
23. Gleason, Henry A.; Cronquist, Arthur. 1991. Manual of vascular plants of
northeastern United States and adjacent Canada. 2nd ed. New York: New
York Botanical Garden. 910 p. [20329]
24. Great Plains Flora Association. 1986. Flora of the Great Plains.
Lawrence, KS: University Press of Kansas. 1392 p. [1603]
25. Harper, K. T.; Sanderson, S. C.; McArthur, E. D. 1992. Riparian ecology
in Zion National Park, Utah. In: Clary, Warren P.; McArthur, E. Durant;
Bedunah, Don; Wambolt, Carl L., compilers. Proceedings--symposium on
ecology and management of riparian shrub communities; 1991 May 29-31;
Sun Valley, ID. Gen. Tech. Rep. INT-289. Ogden, UT: U.S. Department of
Agriculture, Forest Service, Intermountain Research Station: 32-42.
[19092]
26. Harrington, H. D. 1964. Manual of the plants of Colorado. 2d ed.
Chicago: The Swallow Press Inc. 666 p. [6851]
27. Hickman, James C., ed. 1993. The Jepson manual: Higher plants of
California. Berkeley, CA: University of California Press. 1400 p.
[21992]
28. Hitchcock, A. S. 1951. Manual of the grasses of the United States. Misc.
Publ. No. 200. Washington, DC: U.S. Department of Agriculture,
Agricultural Research Administration. 1051 p. [2nd edition revised by
Agnes Chase in two volumes. New York: Dover Publications, Inc.]. [1165]
29. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific
Northwest. Seattle, WA: University of Washington Press. 730 p. [1168]
30. Hoffman, George R. 1985. Germination of herbaceous plants common to
aspen forests of western Colorado. Bulletin of the Torrey Botanical
Club. 112(4): 409-413. [3267]
31. Humphrey, Robert R. 1960. Arizona range grasses: Description--forage
value--management. Tucson, AZ: University of Arizona, Agricultural
Experiment Station. 104 p. [5004]
32. Kearney, Thomas H.; Peebles, Robert H.; Howell, John Thomas; McClintock,
Elizabeth. 1960. Arizona flora. 2d ed. Berkeley, CA: University of
California Press. 1085 p. [6563]
33. Kron, Kathleen A. 1989. The vegetation of Indian Bowl wet prairie and
its adjacent plant communities. I. Description of the vegetation.
Michigan Botanist. 28(4): 179-200. [17358]
34. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation
of the conterminous United States. Special Publication No. 36. New York:
American Geographical Society. 77 p. [1384]
35. Kudish, Michael. 1992. Adirondack upland flora: an ecological
perspective. Saranac, NY: The Chauncy Press. 320 p. [19377]
36. Lackschewitz, Klaus. 1991. Vascular plants of west-central
Montana--identification guidebook. Gen. Tech. Rep. INT-227. Ogden, UT:
U.S. Department of Agriculture, Forest Service, Intermountain Research
Station. 648 p. [13798]
37. Larson, Gary E. 1993. Aquatic and wetland vascular plants of the
Northern Great Plains. Gen. Tech. Rep. RM-238. Fort Collins, CO: U.S.
Department of Agriculture, Forest Service, Rocky Mountain Forest and
Range Experiment Station. 681 p. [22534]
38. Larson, Milo; Moir, W. H. 1987. Forest and woodland habitat types (plant
associations) of northern New Mexico and northern Arizona. 2d ed.
Albuquerque, NM: U.S. Department of Agriculture, Forest Service,
Southwestern Region. 90 p. [8947]
39. Madany, Michael H.; West, Neil E. 1984. Vegetation of two relict mesas
in Zion National Park. Journal of Range Management. 37(5): 456-461.
[6883]
40. Mattson, David John. 1984. Classification and environmental
relationships of wetland vegetation in central Yellowstone National
Park, Wyoming. Moscow, ID: University of Idaho. 409 p. Thesis. [7348]
41. Moir, W. H. 1993. Alpine tundra and coniferous forest. In: Dick-Peddie,
William A., ed. New Mexico vegetation: Past, present, and future.
Albuquerque, NM: University of New Mexico Press: 47-84. [21100]
43. Moir, William H.; Ludwig, John A. 1979. A classification of spruce-fir
and mixed conifer habitat types of Arizona and New Mexico. Res. Pap.
RM-207. Fort Collins, CO: U.S. Department of Agriculture, Forest
Service, Rocky Mountain Forest and Range Experiment Station. 47 p.
[1677]
44. Morgan, M. D. 1969. Ecology of aspen in Gunnison County, Colorado.
American Midland Naturalist. 82(1): 204-228. [15935]
45. Mueggler, W. F. 1985. Vegetation associations. In: DeByle, Norbert V.;
Winokur, Robert P., eds. Aspen: ecology and management in the western
United States. Gen. Tech. Rep. RM-119. Fort Collins, CO: U.S. Department
of Agriculture, Forest Service, Rocky Mountain Forest and Range
Experiment Station: 45-55. [11907]
46. Munz, Philip A. 1973. A California flora and supplement. Berkeley, CA:
University of California Press. 1905 p. [6155]
47. Pfister, Robert Dean. 1972. Vegetation and soils in the subalpine
forests of Utah. Pullman, WA: Washington State University. 98 p.
Dissertation. [16308]
48. Powell, David C. 1988. Aspen community types of the Pike and San Isabel
National Forests in south-central Colorado. R2-ECOL-88-01. Denver, CO:
U.S. Department of Agriculture, Forest Service, Rocky Mountain Region.
254 p. [15285]
49. Radford, Albert E.; Ahles, Harry E.; Bell, C. Ritchie. 1968. Manual of
the vascular flora of the Carolinas. Chapel Hill, NC: The University of
North Carolina Press. 1183 p. [7606]
50. Raunkiaer, C. 1934. The life forms of plants and statistical plant
geography. Oxford: Clarendon Press. 632 p. [2843]
51. Roland, A. E.; Smith, E. C. 1969. The flora of Nova Scotia. Halifax, NS:
Nova Scotia Museum. 746 p. [13158]
52. Rosie, Rhonda. 1991. Range extensions and rare vascular plants from
southeastern Yukon Territory. Canadian Field-Naturalist. 105(3):
315-324. [18205]
53. Schaack, Clark G. 1983. The alpine vascular flora of Arizona. Madrono.
30(4): 79-88. [2069]
54. Smith, D. W.; James, T. D. W. 1978. Changes in the shrub and herb layers
of vegetation after prescribed burning in Populus tremuloides woodland
in southern Ontario. Canadian Journal of Botany. 56: 1792-1797. [16400]
55. Stickney, Peter F. 1989. Seral origin of species originating in northern
Rocky Mountain forests. Unpublished draft on file at: U.S. Department of
Agriculture, Forest Service, Intermountain Research Station, Fire
Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090]
56. Sutton, R. F. 1992. White spruce (Picea glauca [Moench] Voss):
stagnating boreal old-field plantations unresponsive to fertilization
and weed control. Forestry Chronicle. 68(2): 249-258. [18223]
57. U.S. Department of Agriculture, Soil Conservation Service. 1982.
National list of scientific plant names. Vol. 1. List of plant names.
SCS-TP-159. Washington, DC. 416 p. [11573]
58. Voss, Edward G. 1972. Michigan flora. Part I. Gymnosperms and monocots.
Bloomfield Hills, MI: Cranbrook Institute of Science; Ann Arbor, MI:
University of Michigan Herbarium. 488 p. [11471]
59. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry
C., eds. 1987. A Utah flora. Great Basin Naturalist Memoir No. 9. Provo,
UT: Brigham Young University. 894 p. [2944]
60. Wheeler, Gerald A.; Glaser, Paul H.; Gorham, Eville; [and others]. 1983.
Contributions to the Red Lake peatland, northern Minnesota, with special
attention to Carex. American Midland Naturalist. 110(1): 62-96. [18778]
61. Wofford, B. Eugene. 1989. Guide to the vascular plants of the Blue
Ridge. Athens, GA: The University of Georgia Press. 384 p. [12908]
62. Young, Richard P., compiler. 1980. A riparian community classification
study. Cooperative project between Utah State University and the U.S.
Forest Service, Region IV. Final Report. Logan, UT: Utah State
University, Department of Range Science. 77 p. [11734]
63. Zika, Peter F. 1992. Contributions to the alpine flora of the
northeastern United States. Rhodora. 94(877): 15-37. [18123]
64. Bartos, Dale L. 1979. Effects of burning on the aspen ecosystem. In:
Johnson, Kendall L., ed. Wyoming shrublands: Proceedings of the 8th
shrub ecology workshop; 1979 May 30-31; Jackson, WY. Laramie, WY:
University of Wyoming, Division of Range Management, Wyoming Shrub
Ecology Workshop: 47-58. [400]
65. Bartos, Dale L.; Mueggler, Walter F. 1979. Influence of fire on
vegetation production in the aspen ecosystem in western Wyoming. In:
Boyce, Mark S.; Hayden-Wing, Larry D., eds. North American elk, ecology,
behavior and management. Laramie, WY: University of Wyoming: 75-78.
[5101]
66. Seymour, Frank Conkling. 1982. The flora of New England. 2d ed.
Phytologia Memoirs 5. Plainfield, NJ: Harold N. Moldenke and Alma L.
Moldenke. 611 p. [7604]
67. Wright, Henry A.; Bailey, Arthur W. 1980. Fire ecology and prescribed
burning in the Great Plains--a research review. Gen. Tech. Rep. INT-77.
Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain
Forest and Range Experiment Station. 60 p. [2618]
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