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SPECIES:  Menziesia ferruginea
Rusty menziesia branch. Creative Commons image ©2000 Thayne Tuason.

Introductory

SPECIES: Menziesia ferruginea
AUTHORSHIP AND CITATION : Habeck, R. J. 1992. Menziesia ferruginea. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: https://www.fs.usda.gov/database/feis/plants/shrub/menfer/all.html []. Revisons : On 14 December 2017, the common name of this species was changed in FEIS from: menziesia to: rusty menziesia. The photo was also added.
ABBREVIATION : MENFER SYNONYMS : Menziesia glabella (Gray) Peck NRCS PLANT CODE : MEFE COMMON NAMES : rusty menziesia false huckleberry fool's huckleberry false azalea menziesia mock azalea smooth menziesia Pacific menziesia skunkbrush rustyleaf TAXONOMY : The currently accepted scientific name of rusty menziesia is Menziesia ferruginea (Smith) [26]. In existence are two varieties distinguished by morphological variations and geographic location [25,26,32,33]: M. ferruginea var. ferruginea M. ferruginea var. glabella (Gray) Peck LIFE FORM : Shrub FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY

DISTRIBUTION AND OCCURRENCE

SPECIES: Menziesia ferruginea
GENERAL DISTRIBUTION : Menziesia ferruginea var. ferruginea is found north along the Alaskan Coast, and south along the Pacific Coast to northern California, inland across the Cascades and Rocky Mountains to Wyoming [25,33]. Menziesia ferruginea var. glabella is distributed from British Columbia to Alberta, south to Montana, Wyoming, Idaho, eastern Washington, and Oregon; down the Columbia River to Mt. Hood and Mt. Adams, where the two varieties freely interbreed [25,27]. ECOSYSTEMS :    FRES20  Douglas-fir    FRES22  Western white pine    FRES23  Fir - spruce    FRES24  Hemlock - Sitka spruce    FRES25  Larch STATES :      AK  CA  ID  MT  OR  WA  WY  AB  BC BLM PHYSIOGRAPHIC REGIONS :    1  Northern Pacific Border    2  Cascade Mountains    4  Sierra Mountains    5  Columbia Plateau    8  Northern Rocky Mountains    9  Middle Rocky Mountains KUCHLER PLANT ASSOCIATIONS :    K001  Spruce - cedar - hemlock forest    K002  Cedar - hemlock - Douglas-fir forest    K003  Silver fir - Douglas-fir forest    K004  Fir - hemlock forest    K012  Douglas-fir forest    K013  Cedar - hemlock - pine forest    K014  Grand fir - Douglas-fir forest    K015  Western spruce - fir forest SAF COVER TYPES :    205  Mountain hemlock    206  Engelmann spruce - subalpine fir    210  Interior Douglas-fir    212  Western larch    213  Grand fir    215  Western white pine SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Rusty menziesia is an understory dominant on moist, wooded north- and east-facing slopes of the montane to upper subalpine zones.  Overstory components mostly include subalpine fir (Abies lasiocarpa), Pacific silver fir (A. amabilis), western redcedar (Thuja plicata), mountain hemlock (Tsuga mertensiana), western hemlock (T. heterophylla), and Engelmann spruce (Picea engelmannii) [25].  Rusty menziesia's associates generally include blue huckleberry (Vaccinium globulare), smooth woodrush (Luzula hitchcockii), and on higher sites, white rhododendron (Rhododendron albiflorum) [33].  In Alaska, rusty menziesia is a common understory shrub in the coastal spruce-hemlock forest, often under a dense canopy.  It also grows in the southern part of the boreal forest in white spruce (Picea glauca) and white spruce-paper birch (Betula papyrifera) stands [53]. Publications listing rusty menziesia as an indicator or dominant species in habitat types (hts), community types (cts), or plant associations (pas) are presented below: Area                       Classification       Authority   AK                        forest (cts)        Reynolds 1990 c ID                        forest (hts)        Steele & others  1981 n ID                        forest (hts)        Cooper & others  1991 e ID and w WY               forest (hts)        Steele & others  1983   MT                        forest (hts)        Pfister & others 1977   OR: Mt. Hood &       Willamette N.F.       forest (pas)        Hemstrom & others 1982   OR: Wallowa-Whitman       National Forest       forest (pas)        Johnson & Simon   1987   WA: Mount Rainier N.P.    forest (pas)        Franklin & others 1988   WA: Gifford Pinchot       National Forest       forest (pas)        Brockway & others 1983

MANAGEMENT CONSIDERATIONS

SPECIES: Menziesia ferruginea
IMPORTANCE TO LIVESTOCK AND WILDLIFE : Rusty menziesia's value to wildlife is primarily as food.  In Idaho and Montana, elk used rusty menziesia in considerable amounts in July, and to a lesser extent in October.  Elk rarely eat rusty menziesia when it occurs with more palatable browse species.  Menziesia filled a browse requirement during July when elk were primarily on an herbaceous diet in western Montana and Idaho [14,54].  Mule deer use rusty menziesia moderately in summer and fall in the western states [31].  Rusty menziesia accounted for over 15 percent of the summer moose diet in the spruce-fir cover types of Jackson Hole, Wyoming [28].  Alaskan spruce grouse use small amounts of rusty menziesia capsules in their October diet [15].  Following a burn treatment in western Montana, rusty menziesia came back as an important source of fruits and seeds used by rodents such as deer mice, chipmunks, and voles [21]. PALATABILITY : The palatability of rusty menziesia is generally rated poor.  This shrub provides fair summer browse for deer and elk [35].  Rusty menziesia's palatability was considered very low for moose in Jackson Hole, Wyoming [23]. NUTRITIONAL VALUE : Nutrient value of rusty menziesia is considered poor to medium [20]. Rusty menziesia contains toxic resinoids that are known to cause livestock losses when leaves and roots are eaten [43]. Mean values of percent composition of macro- and micronutrients for rusty menziesia are as follows [22]:                     MACRO                              MICRO              (percent dry weight)                 (ppm dry weight) ---------------------------------------------------------------------           N   P    K    Ca   Mg   Na           Cu     Mn    Fe   Zn --------------------------------------------------------------------- Leaves  2.48 0.24 1.19 0.62 0.40 0.47         10.0  16,656 62.0 86.6 Stems   1.18 0.15 0.85 0.35 0.17 0.09         11.8  7139.2 22.8 40.5 COVER VALUE : Rusty menziesia provides shelter and cover for a variety of wildlife species. Dense thickets provide good cover for many small birds and mammals. VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : Rusty menziesia is a desirable ornamental due its crimson-orange autumn foliage [26]. OTHER MANAGEMENT CONSIDERATIONS : Rusty menziesia is mostly threatened by logging activities.  Following clearcutting of Pacific silver fir and mountain hemlock, rusty menziesia exhibits a difficult time regenerating [6].  In northwestern Montana, rusty menziesia decreased considerably from its high abundance following a clearcut in an old-growth stand [2].  Soil layers restricting root growth and high water tables are common enough that compaction or erosion-causing activities should be carefully controlled [6]. There appears to be little destruction of mature rusty menziesia plants in spruce-fir forests when selective logging is done during the winter [9]. If rusty menziesia is a major component of the mature forest understory, it will continue to dominate the postharvest plant community [10]. Mechanical site preparation:  Coates [8] found that recovery of rusty menziesia on Engelmann spruce and subalpine fir sites after clipping and mechanical scarification was very limited and confined to the production of new shoots from buds on old stems rather than from basal sprouts or suckers.  Two seasons after cutting, new shoots had not reached the height of precut stems, and did not overtop spruce or pine seedlings planted at the time of cutting. Rusty menziesia control by Esteron brush killer was best achieved when this herbicide was mixed with diesel and sprayed by helicopter from 150 to 200 feet (46-61 m) high [40].  After 2 years, Tordon 101 and Banvel 720 killed 100 percent of rusty menziesia when sprayed on leaf foliage [39].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Menziesia ferruginea
GENERAL BOTANICAL CHARACTERISTICS : Rusty menziesia is an erect, deciduous shrub 3.3 to 6.6 feet (1-2 m) tall. Ovate-elliptic to elliptic-obovate leaves are 1.5 to 2.5 inches (4-6 cm) long and exude a skunky odor when crushed.  Rusty menziesia has scaly bark on older branches, while the young twigs are covered with fine hair [20]. The fruits are oval capsules containing many seeds 0.20 to 0.28 inch (5-7 mm) long.  The leaves and stems of this shrub are sticky to the touch [25,26,44]. Rusty menziesia decreases in height with increasing elevation.  This would imply that rusty menziesia's height is generally at its maximum on western hemlock sites, and at its minimum on mountain hemlock and subalpine fir sites [34]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Seed production and dispersal:  Numerous small seeds are produced in capsules which dehisce naturally at maturity.  Seeds are wind or gravity dispersed. Vegetative Reproduction:  Rusty menziesia sprouts from root crowns and adventitious stem buds after destruction of aboveground parts.  When covered with ash from Mt. St. Helen's, rusty menziesia produced adventitious roots [3].  This plant also appears to be capable of layering.  Layering is an effective means of growth and expansion at high elevations where heavy snowpacks press rusty menziesia branches close to the ground [10].  On steep slopes, rusty menziesia tends to grow downhill with the tips of its branches ascending [35]. SITE CHARACTERISTICS : Rusty menziesia is a common shade-tolerant shrub, found predominantly on steep, northerly exposures with abundant moisture [17,20,25,26]. Rusty menziesia generally occurs from 3,300 to 7,500 feet (1,000-2,300 m) in elevation on 15 to 60 percent slopes [11,29,50].  Absence from extreme northern distributions may imply a limitation to cold temperatures. Rusty menziesia may be restricted to areas with adequate snow cover, which protects plants from winter desiccation [25].  In Oregon, rusty menziesia is typically found on high-elevation sites where frequent frost, heavy snowpacks, and short growing seasons are common [6]. The presence of rusty menziesia has been reported as being a particularly good indicator of site potential [13].  Rusty menziesia often grows under dense canopies, in openings, and on cut-over forest land, especially on well-drained slopes [53].  Mueggler [42] found the distribution of rusty menziesia in cedar-hemlock stands to be positively associated with low soil potassium content, and high organic matter content (5.6 to 8.0 percent). Soils:  Soil depth usually ranges from 17 to 70 inches (43-178 cm). Soils are generally derived from loam to sandy-loam pumice layers to a depth of 15 to 30 inches (38-76 cm) [6,11,24].  In central Idaho, soil acidity ranges from pH 5.1 to 6.2; litter depth can average at least 3.5 inches (9 cm) [6,11].  Effective rooting depth in Oregon is approximately 46 inches (117 cm) [6]. SUCCESSIONAL STATUS : Rusty menziesia is most always associated with cold, wet habitat types. Rusty menziesia is very shade tolerant.  It can persist in heavily shaded forests where light levels are typically 5 percent of full sunlight. Alaback [1] reported that rusty menziesia decreased in biomass productivity in Alaskan spruce-hemlock forests over 30 years of age.  Rusty menziesia biomass was lowest in stands 31 to 130 years old, but after this period increased.  In western Montana, burning increased rusty menziesia fourfold from immature stands (less than 90 years) to old stands (greater than 150 years) [2]. SEASONAL DEVELOPMENT : Phenology:  Rusty menziesia flowers from late May through July; capsules mature in July and August [53].  Leaves expand rapidly in early June at the time flowers open and begin to senesce in mid-August and September at high elevations in southern British Columbia [10].  Rusty menziesia has been found to live at least 30 years in northern forests [1].

FIRE ECOLOGY

SPECIES: Menziesia ferruginea
FIRE ECOLOGY OR ADAPTATIONS : Rusty menziesia is a fire-sensitive species [51].  It responds to fire by sprouting from the root crown [38].  Since rusty menziesia is indicative of cool, moist sites, fire plays a minor role in its development.  Barrett and Arno [4] found that the subalpine fir/rusty menziesia cover type in the Selway-Bitterroot Wilderness, Idaho, had a mean fire interval of 174 to 181 years, achieving lethal/nonuniform fire behavior. FIRE REGIMES : Find fire regime information for the plant communities in which this species may occur by entering the species name in the FEIS home page under "Find Fire Regimes". POSTFIRE REGENERATION STRATEGY :    survivor species; on-site surviving root crown or caudex

FIRE EFFECTS

SPECIES: Menziesia ferruginea
IMMEDIATE FIRE EFFECT ON PLANT : Rusty menziesia is very susceptible to fire-kill.  Moderate to severe fires reduce survival and slow redevelopment [44].  Wildfires apparently removed rusty menziesia totally from experimental plots in Washington and Oregon for up to 16 years [58]. PLANT RESPONSE TO FIRE : In western Montana, Stickney [51] found that postfire survival of rusty menziesia was poor; rusty menziesia was very slow to redevelop after fire, especially in areas that had been intensely burned.  The average percent cover of rusty menziesia doubled from unburned plots to lightly burned plots, and tripled from moderately burned plots in western Montana [48]. Another study found rusty menziesia to increase its density of aboveground parts the first year after burning [37].  Model projections predict that rusty menziesia will reach a height of 2.4 feet (.73 m) 2 years after a burn, and that the average height of rusty menziesia will peak at 5.1 to 5.6 feet (1.5-1.7 m) 20 to 25 years after burning [34].  Mueggler [42], however, found no significant difference in cover and frequency between burned and unburned sites in northern Idaho. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : The following Research Project Summaries provide information on prescribed fire use and postfire response of plant community species including rusty menziesia: Hamilton's Research Papers (Hamilton 2006a, Hamilton 2006b) also provide information on prescribed fire and postfire response of plant species, including rusty menziesia, that was not available when this species review was originally written. FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Menziesia ferruginea
REFERENCES :  1.  Alaback, Paul B. 1982. Dynamics of understory biomass in Sitka        spruce-western hemlock forests of southeast Alaska. Ecology. 63(6):        1932-1948.  [7305]  2.  Antos, J. A.; Habeck, J. R. 1981. Successional development in Abies        grandis (Dougl.) Forbes forests in the Swan Valley, western Montana.        Northwest Science. 55(1): 26-39.  [12445]  3.  Antos, Joseph A.; Zobel, Donald B. 1985. Plant form, developmental        plasticity and survival following burial by volcanic tephra. Canadian        Journal of Botany. 63: 2083-2090.  [12553]  4.  Barrett, Stephen W.; Arno, Stephen F. 1991. Classifying fire regimes and        defining their topographic controls in the Selway-Bitterroot Wilderness.        In: Andrews, Patricia L.; Potts, Donald F., eds. Proceedings, 11th        annual conference on fire and forest meteorology; 1991 April 16-19;        Missoula, MT. SAF Publication 91-04. Bethesda, MD: Society of American        Foresters: 299-307.  [16179]  5.  Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals,        reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's        associations for the eleven western states. Tech. Note 301. Denver, CO:        U.S. Department of the Interior, Bureau of Land Management. 169 p.        [434]  6.  Brockway, Dale G.; Topik, Christopher; Hemstrom, Miles A.; Emmingham,        William H. 1985. Plant association and management guide for the Pacific        silver fir zone: Gifford Pinchot National Forest. R6-Ecol-130a.        Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific        Northwest Region. 122 p.  [525]  7.  Cates, Rex G.; Orians, Gordon H. 1975. Successional status and the        palatability of plants to generalized herbivores. Ecology. 56: 410-418.        [15989]  8.  Coates, K. David. 1987. Effects of shrubs and herbs on conifer        regeneration and microclimate in the Rhododendron-Vaccinium-Menziesia        community of south-central BC. Vancouver, BC: University of British        Columbia. Thesis. Abstract.  [17445]  9.  Coates, D.; Haeussler, S. 1986. A preliminary guide to the response of        major species of competing vegetation to silvicultural treatments.        Victoria, BC: Ministry of Forests, Information Services Branch; Land        Management Handbook Number 9. 88 p.  [17453] 10.  Comeau, Philip G.; Watts, Susan B.; Caza, Caroline L.; [and others].        1989. Autecology, biology, competitive status and response to treatment        of seven southern interior weed species. FRDA Report 093; ISSN 0835        0572. Victoria, BC: BC Ministry of Forests, Research Branch. 46 p.        [9471] 11.  Cooper, Stephen V.; Neiman, Kenneth E.; Roberts, David W. 1991. (Rev.)        Forest habitat types of northern Idaho: a second approximation. Gen.        Tech. Rep. 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FEIS Home Page
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