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SPECIES:  Ribes lacustre
Prickly currrant. Image by Rob Routledge, Sault College, Bugwood.org.

Introductory

SPECIES: Ribes lacustre
AUTHORSHIP AND CITATION: Carey, Jennifer H. 1995. Ribes lacustre. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: https://www.fs.usda.gov/database/feis/plants/shrub/riblac/all.html []. Revisions: On 21 August 2018, the common name of this species was changed in FEIS from: bristly black currant to: prickly currant. Images were also added. ABBREVIATION: RIBLAC SYNONYMS: NO-ENTRY NRCS PLANT CODE: RILA COMMON NAMES: prickly currant bristly black currant swamp currant TAXONOMY: The scientific name of prickly currant is Ribes lacustre (Pers.) Poir. (Grossulariaceae) [25,33,34,35,59]. There are no currently accepted infrataxa. LIFE FORM: Shrub FEDERAL LEGAL STATUS: No special status OTHER STATUS: NO-ENTRY

DISTRIBUTION AND OCCURRENCE

SPECIES: Ribes lacustre
GENERAL DISTRIBUTION: Prickly currant occurs throughout Canada from Newfoundland to Yukon Territory and in Alaska [35,54,59].  It extends south into the continental United States in the Coast and Cascade ranges to northern California, in the Rocky Mountains to central Colorado and northern Utah, in the Great Lake States, and in the Appalachian Mountains to West Virginia [8,25,33,65,72].  In the northern Great Plains region, prickly currant occurs in the Black Hills [26].
Distribution of prickly currant. Map courtesy of USDA, NRCS. 2018. The PLANTS Database. National Plant Data Team, Greensboro, NC [2018, August 21] [67].
ECOSYSTEMS: 
   FRES11  Spruce-fir
   FRES18  Maple-beech-birch
   FRES19  Aspen-birch
   FRES20  Douglas-fir
   FRES22  Western white pine
   FRES23  Fir-spruce
   FRES24  Hemlock-Sitka spruce
   FRES25  Larch
   FRES26  Lodgepole pine
   FRES28  Western hardwoods


STATES: 
     AK  CA  CO  CT  ID  ME  MA  MI  MN  MT
     NH  OH  OR  PA  SD  UT  VT  VA  WA  WV
     WY  AB  BC  MB  NB  NF  NT  ON  PE  PQ
     SK  YT



BLM PHYSIOGRAPHIC REGIONS: 
    1  Northern Pacific Border
    2  Cascade Mountains
    5  Columbia Plateau
    8  Northern Rocky Mountains
    9  Middle Rocky Mountains
   10  Wyoming Basin
   11  Southern Rocky Mountains
   15  Black Hills Uplift
   16  Upper Missouri Basin and Broken Lands


KUCHLER PLANT ASSOCIATIONS: 
   K001  Spruce-cedar-hemlock forest
   K003  Silver fir-Douglas-fir forest
   K004  Fir-hemlock forest
   K007  Red fir forest
   K008  Lodgepole pine-subalpine forest
   K012  Douglas-fir forest
   K013  Cedar-hemlock-pine forest
   K014  Grand fir-Douglas-fir forest
   K015  Western spruce-fir forest
   K025  Alder-ash forest
   K093  Great Lakes spruce-fir forest
   K096  Northeastern spruce-fir forest
   K106  Northern hardwoods


SAF COVER TYPES: 
    60  Beech-sugar maple
   107  White spruce
   201  White spruce
   205  Mountain hemlock
   206  Engelmann spruce-subalpine fir
   207  Red fir
   208  Whitebark pine
   210  Interior Douglas-fir
   211  White fir
   212  Western larch
   213  Grand fir
   215  Western white pine
   218  Lodgepole pine
   221  Red alder
   222  Black cottonwood-willow
   223  Sitka spruce
   227  Western redcedar-western hemlock


SRM (RANGELAND) COVER TYPES: 
   422  Riparian


HABITAT TYPES AND PLANT COMMUNITIES: 
Prickly currant occurs in woods, forests, and shrublands.  It is a
common but not abundant understory species [16,27,32].

Prickly currant occurs in the understory of subalpine forests in
Wyoming with gooseberry currant (Ribes montigenum), sidebells
wintergreen (Orthilia secunda), heartleaf arnica (Arnica cordifolia),
and fireweed (Epilobium angustifolium) [9].

In the cedar (Thuja spp.)-hemlock (Tsuga spp.) zone of northwestern
British Columbia, prickly currant occurs with devil's club
(Oplopanax horridus), leafy moss (Mnium spp.), oak fern (Gymnocarpium
dryopteris), Schreber's moss (Pleurozium schreberi), common ladyfern
(Athyrium filix-femina), and horsetail (Equisetum spp.) [28].  
Prickly currant occurs in the oak fern, devil's club, horsetail, and
queencup beadlily (Clintonia uniflora) series of the wet, cool subboreal
spruce (Picea spp.) forest zone in British Columbia [32].

Prickly currant occurs with Rocky Mountain maple (Acer glabrum),
Utah honeysuckle (Lonicera utahensis), blue huckleberry (Vaccinium
membranaceum), and western meadowrue (Thalictrum occidentale) in the
understory of a virgin grand fir (Abies grandis) forest in northern
Idaho [36].

In spruce forests in Alberta, prickly currant occurs with other
mesophytic species including twinberry honeysuckle (Lonicera
involucrata), highbush cranberry (Viburnum edule), tall bluebells
(Mertensia paniculata), wild sarsaparilla (Aralia nudicaulis), bluejoint
reedgrass (Calamagrostis canadensis), oak fern, stiff clubmoss
(Lycopodium amnotinum), and claspleaf twistedstalk (Streptopus
amplexifolius) [41].

Prickly currant occurs in riparian woodlands and shrublands.  It
occurs in a thinleaf alder (Alnus incana ssp. tenuifolia) riparian
dominance type in east-central Oregon [50].  In the Klamath Mountains of
northern California, prickly currant borders streams with thinleaf
alder, California mountain-ash (Sorbus californica), and Scouler willow
(Salix scouleriana), and it occurs in thickets with thinleaf alder,
Sitka alder (Alnus viridis ssp. sinuata), red-osier dogwood (Cornus
sericea), red elderberry (Sambucus racemosa ssp. pubens), and cascara
(Rhamnus purshiana) [56].

Clearcuts in the Olympic Mountains in Washington, dominated by
oceanspray (Holodiscus discolor) and thimbleberry (Rubus parviflorus),
include common snowberry (Symphoricarpos albus) and prickly
currant [23].

MANAGEMENT CONSIDERATIONS

SPECIES: Ribes lacustre
IMPORTANCE TO LIVESTOCK AND WILDLIFE: Prickly currant berries are eaten by rodents, bears, and birds [38].  In southwestern Alberta grizzly bears feed on prickly currant berries in late summer and early autumn [31].  Elk, mule deer, white-tailed deer, and mountain goats eat prickly currant foliage [11,18,55,58,71].  In Montana prickly currant was 1 percent of elk diet in early summer and 3 percent in late summer [18]. PALATABILITY: Prickly currant browse is moderately palatable to livestock, elk, and deer [17,38].  The berries are edible but somewhat disagreeable tasting [35,38,59]. NUTRITIONAL VALUE: Prickly currant berries collected in the summer in northern Ontario consisted of 79.68 percent moisture, 20 percent dry matter, 0.28 percent fat, 1.46 percent protein, and 5.66 percent soluble carbohydrate based on fresh fruit weight.  Fresh prickly currant fruit contains slightly more than 30 kilocalories per 100 grams [69]. Nutritional values of prickly currant berries collected in late summer in southeastern Washington, based on dry weight, were 2.94 percent protein, 4.09 percent lipid, 3.96 percent neutral detergent fiber, 4.56 percent ash, 0.282 percent calcium, 0.066 percent magnesium, 0.17 percent phosphorus, and 1.887 percent potassium [52]. Norton and others [46] analyzed the nutritional value of dried prickly currant berries stored for 1 year.  Traditional Native American drying and storing methods were used.  One gram dry weight contained on average 3.19 milligrams calcium, 0.05 milligram iron, 0.94 milligram magnesium, 0.02 milligram zinc, and 3.33 milligrams ascorbic acid [46]. COVER VALUE: In Wyoming, prickly currant is considered poor cover for elk and pronghorn and fair cover for mule deer and white-tailed deer.  It is good cover for upland game birds, small nongame birds, and small mammals [17]. VALUE FOR REHABILITATION OF DISTURBED SITES: NO-ENTRY OTHER USES AND VALUES: Native Americans in the Pacific Northwest ate prickly currant berries in historic times [46]. OTHER MANAGEMENT CONSIDERATIONS: Prickly currant is an alternate host for white pine blister rust (Cronartium ribicola) which infests five-needled pines.  Because of its association with the rust, prickly currant has been a target of various eradication studies.  Prickly currant is highly resistant to chemical injury [48].  Efforts to eradicate Ribes spp. have been unsuccessful and have not resulted in decreased rust infection.  Only a few Ribes bushes per acre are sufficient to perpetuate blister rust [29]. Although prickly currant establishes on scarified ground after tree harvest, it does not impede conifer seedling establishment [61]. Grazing by wild ungulates prevents development of prickly currant in clearcuts.  Eleven years after clearcutting, burned and unburned grazed sites averaged 0.3 percent cover prickly currant.  Ungrazed burned clearcuts averaged 4.6 percent cover and ungrazed unburned clearcuts averaged 9.8 percent cover [19].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Ribes lacustre
GENERAL BOTANICAL CHARACTERISTICS: Prickly currant is a native, deciduous shrub that grows 3 to 4 feet (1-1.2 m) tall [44,59,72].  In sunlight prickly currant grows erect, but in shade, branches are often reclining or trailing [44]. Prickly currant has prickly stems and nodal spines.  The drooping raceme has 5 to 15 flowers [33].  The berries average 0.34 inch (8.6 mm) in diameter and contain an average of 16.5 small seeds [52]. The root systems of Ribes spp. consist of shallow roots radiating from a central root crown [47].  Prickly currant roots are very shallow, especially on moist sites, but lateral spread can be extensive [48]. Some sources report that prickly currant is rhizomatous [22,48] while others report that it is nonrhizomatous [61].  Prickly currant partially buried by volcanic ejecta from Mount St. Helens in southern Washington had not developed rhizomes but had a well-developed adventitious root system [3]. RAUNKIAER LIFE FORM: Phanerophyte REGENERATION PROCESSES: Prickly currant regenerates primarily from seed.  Plants first begin producing seeds when 3 to 5 years old.  An average bush produces 50 to 75 berries.  Good crops occur in 2- to 3-year intervals.  Some seeds are dispersed by animals, but many berries fall to the ground beneath the parent plant [44]. Stratification is usually required to break the dormancy in prickly currant seed.  Seeds stored at 32 degrees Fahrenheit (0 deg C) for 120 to 200 days had 48 percent germination in sand moistened with nutrient solution.  Alternating diurnal temperatures (77 degrees Fahrenheit [25 deg C] and 41 or 50 degrees Fahrenheit [5 or 10 deg C]) results in some germination without prior stratification.  Scarification enhances germination.  A five-minute soak in 2 to 10 percent sulfuric acid solution improved germination [51]. Mineral soil is the best seed bed.  Prickly currant establishes on well scarified sites [44,61]. Prickly currant seeds have long-term viability.  They accumulate in the organic mantle and mineral soil over time.  The mineral soil seedbank in mature forests in west-central Idaho contained 51 viable prickly currant and sticky currant (Ribes viscosissimum) seeds per square foot (567/sq m).  Over 80 percent of the viable seeds were found in the top 2 inches (5 cm) of mineral soil.  The two Ribes spp. were combined in the data because seedlings could not be distinguished in the greenhouse [39]. Prickly currant regenerates vegetatively [44,48].  Prickly currant stems in contact with soil produce adventitious roots [3,27,48]. In southern Washington, prickly currant that was partially buried for 1 year by 2 to 8 inches (5-20 cm) of volcanic ejecta had two to five adventitious roots per centimeter of stem.  Maximum adventitious root length was 10 inches (25 cm) [3].  Offord and others [48] observed rhizomes in prickly currant near Mount Hood in northwestern Oregon.  Some sources [22,43,47] suggest that prickly currant sprouts from the root crown, but definitive documentation was not found in the literature. SITE CHARACTERISTICS: Prickly currant occurs in moist woods and forests, ravines, shrub thickets, meadow margins, swamps, rock crevices, seepage areas, along avalanche chutes, on streambanks, and on steep slopes [4,27,33,34,59]. On Big Snowy Peak in central Montana, prickly currant grows in crevices of limestone outcrops and cliffs [5]. Prickly currant occurs in cool, moist climates.  It occurs from 7,700 to 10,500 feet (2,300-3,200 m) in Utah, 7,000 to 11,400 feet (2,100-3,500 m) in Colorado, 5,500 to 10,700 feet (1,700-3,300 m) in Wyoming, and 3,000 to 8,700 feet (900-2,700 m) in Montana [17].  At the southern extremes of its range (California, Utah, Colorado, and West Virginia), prickly currant occurs in cool high-elevation forests [1,56,65].  It is found more often on northerly and easterly exposures than southerly or westerly exposures [44]. Prickly currant occurs on moist, nutrient-rich sites.  In 91- to 160-year-old lodgepole pine (Pinus contorta) stands and 60- to 160-year-old white spruce (Picea glauca) stands in west-central Alberta, prickly currant had significantly (P<0.025) higher canopy cover on sites with high site index (indicative of higher productivity) than low site index [66].  In the Prince Rupert Forest Region of British Columbia, prickly currant occurs on mesic to subhydric sites with high nutrient status (permesotrophic to subeutrophic) [73].  It occurs in both acidic and basic soils [1,5]. SUCCESSIONAL STATUS: Prickly currant is moderately shade tolerant [32,44] but grows most vigorously in canopy openings [27].  It establishes in partial shade or full sun after disturbance and then persists in the understory of closed canopy forests and woods [10,44,61].  Seedlings are suppressed on sites with more than 75 percent of full shade [44].  Prickly currant established on a 1-year-old mudflow surface formed by the volcanic eruption of Mount St. Helens.  The mudflow was reworked nonorganic substrate with no shallowly buried soil [30] Dense thickets of prickly currant are uncommon.  Prickly currant canopy cover reported in the literature is generally less than 10 percent and commonly less than 1 percent [16,27,32]. Prickly currant response to timber harvest is an indicator of its successional status.  Although prickly currant generally increases after clearcutting [2,20,23,32], its response is varied.  In western Montana prickly currant cover averaged 1.5 percent in 7- to 16-year-old clearcuts but was only a trace in uncut stands [2].  In northern British Columbia, prickly currant increased in abundance on alluvial and poorly drained sites following clearcutting [20]. Prickly currant is a principal understory species on clearcuts in northwestern Washington [23].  In subalpine forests in central Colorado, pre- and postlogging prickly currant canopy cover was not significantly (P<0.05) different [13].  In northwestern Montana, prickly currant response to clearcutting depended on habitat type.  In the subalpine fir/queencup beadlily habitat type, prickly currant canopy cover was lower on burn sites, clearcut sites, and avalanche chutes than on old growth sites; in the more mesic subalpine fir/menziesia (Menziesia spp.)  habitat type, prickly currant canopy cover was higher after these disturbances than on old-growth sites [74].   SEASONAL DEVELOPMENT: Prickly currant flowers from May to June, and berries ripen in August.  Seeds germinate in the spring [51].  Prickly currant does not always produce fruit in spruce-fir (Abies spp.) forests of Canada because of the short growing season [27].

FIRE ECOLOGY

SPECIES: Ribes lacustre
FIRE ECOLOGY OR ADAPTATIONS: Prickly currant occurs in many forest types, such as grand fir and Engelmann spruce (Picea engelmannii)-subalpine fir (Abies lasiocarpa), that are characterized by long fire-free intervals punctuated by severe, stand-replacing fires [22,77].  The ability of prickly currant to regenerate after fire from long-lived seed stored in soil or from off-site sources makes prickly currant fairly resilient to stand-replacing fire [89].  Low-severity fire is probably more favorable to Ribes spp. than severe fire [78]. After fire prickly currant canopy cover increases slowly, reaching a maximum density in several to many decades [15,42,76].  Prickly currant cover was reported for sites with a history of fire in the southern boreal forest region of Quebec.  Postfire ages of sites ranged from 26 to 230 years.  Prickly currant was present on all sites at low cover.  The highest prickly currant cover recorded (2.2 percent) occurred on the site with a postfire age of 74 [15].  The percent prickly currant cover follows:                                      Years since fire                   26  46 74 120 143 167 174 230 Canopy cover      0.1    0.2 2.2 1.0 0.3 0.1 0.1 0.1 In north-central Idaho, the vegetation was measured for 36 clearcut and broadcast burned sites and 7 old-growth sites.  Postfire ages of burned sites ranged from 1 to 23 years.  Prickly currant average percent canopy volume (percent of total volume occupied in a 1x1x3 m space) and average height by postfire year follow [75]:                                       Years since fire                          1       3  8 12 23  old growth Avg. % canopy volume  0.1  0.1  0.3 0.8    trace Average height (cm)      11  24  45  65 43      25 POSTFIRE REGENERATION STRATEGY:    Ground residual colonizer (on-site, initial community)    Secondary colonizer - off-site seed FIRE REGIMES: Find fire regime information for the plant communities in which this species may occur by entering the species name in the FEIS home page under "Find Fire Regimes".

FIRE EFFECTS

SPECIES: Ribes lacustre
IMMEDIATE FIRE EFFECT ON PLANT: Fire that burns the organic soil probably kills prickly currant. Noste and Bushey [47] report that fire that removes the organic soil layer will likely kill the shallow root systems of most Ribes spp. Prickly currant was nearly absent from study sites after a severe prescribed fire in central Idaho (see Fire Case Studies) [42]. Low-severity fire probably top-kills prickly currant. Prickly currant seeds contained in the organic mantle are destroyed by severe fire [44].  However, seeds buried in mineral soil probably survive most fire. PLANT RESPONSE TO FIRE: Numerous secondary sources suggest that prickly currant sprouts from the root crown and rhizomes after top-kill by fire [7,12,22,43, 47,77].  However, primary documentation of prickly currant sprouting from the root crown or rhizomes after fire was not found in the literature. Prickly currant was present in the postfire ravine vegetation 1 year after a mid-summer wildfire in western Montana.  Although the fire was generally severe, some ravine sites did not burn severely because of the higher moisture content of plant material and soil.  Prickly currant plants recovered quickly in low-severity burn patches; the mechanism of recovery (sprouting from rootstocks, rhizomes, or adventitious stem buds) was not reported.  Maximum prickly currant postfire canopy cover was 2.5 percent 1 year after fire and 6.3 percent 2 years after fire [37]. Prickly currant colonizes burned sites via long-lived buried seed and/or seed carried on to the site by animals [64].  Regeneration success after fire depends on site conditions and fire severity.  Severe fire substantially reduces or delays prickly currant establishment [42].  Prickly currant seedlings were present on 1 of 18 burn sites 1 year after the Sundance wildfire in northern Idaho.  The fire consumed the overstory, litter and duff; fire intensity ranged from 3,400 to 22,500 British thermal units per second per foot [63]. According to Steele and Geier-Hayes [61], prickly currant and sticky currant are common on scarified portions of past pile-and-burn sites but rare on severely burned areas.  In western Montana black currant was 25.6 percent less frequent on burned slash pile sites than on adjacent unburned scarified clearcut sites.  Postfire age of sampled burned sites averaged 8.8 years [70]. Optimal conditions for prickly currant establishment may not occur until several years after fire.  In broadcast-burned clearcuts in north-central Idaho, prickly currant occurred in only 1 of 10 microplots in 1-year-old burns but was found on seven of nine microplots in 3-year-old burns [75].  Severe hot or dry soil conditions associated with some burns may discourage immediate colonization by prickly currant.  Lyon [42] suggests that optimal environmental conditions for prickly currant establishment may not occur on some sites until other vegetation is established. Prickly currant recovery was slow after prescribed broadcast fires in clearcut western larch (Larix occidentalis)-Douglas-fir (Pseudotsuga menziesii) forests in northwestern Montana.  Prickly currant disappeared from the most severely burned site (E-8), but appeared within 8 years on three sites on which it had not been present in the prefire vegetation.  Prickly currant pre- and postfire canopy cover follows [62]:                               Year since fire Site    prefire     1     2     3     4     5     6     7     8   postfire         % cover                                                   duff (cm) N-6        -        -     -     1     1     1     -     -     -     7.0 N-8        -        -     -     -     -     1     -     1     3     6.0 E-8        1        -     -     -     -     -     -     -     -     6.6 S-2        1        -     -     -     -     -     1     -     -     * W-3        -        -     -     -     -     -     -     -     1     4.5 *Duff depth was not reported for the S-2 burn site. Literature which contains fire response information for Ribes spp. in general should be viewed with caution because of substantial differences between co-occurring species.  In the above-mentioned study [62], sticky currant rapidly colonized some burned sites, reaching a maximum canopy cover of 11 percent after 4 years on a site from which it was absent before fire [62].  Lyon [42] also documented rapid sticky currant recovery after fire (see Fire Case Studies).  Mueggler [45] combined Ribes spp. when he reported they were considerably more frequent on burn sites than unburned sites in grand fir and western redcedar (Thuja plicata) forest associations in northern Idaho.  Prickly currant and sticky currant co-occur in these forests [14], and it is possible that only sticky currant was doing well on burned sites in the first several postfire years. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE: For further information on prickly currant response to fire, see Fire Case Studies.  The Research Project Summary and Research Papers (Hamilton 2006a, Hamilton 2006b) of Hamilton's studies and Lyon's Research Paper (Lyon 1966) also provide information on prescribed fire and postfire response of plant community species, including prickly currant. FIRE MANAGEMENT CONSIDERATIONS: Nutrient content of shrub species was estimated for seven silvicultural treatments including burning in the Coram Experimental Forest in northwestern Montana [60].  The elemental content of prickly currant branches and leaves collected in July and August is presented [60].

FIRE CASE STUDY

SPECIES: Ribes lacustre
FIRE CASE STUDY CITATION: Carey, Jennifer H., compiler. 1995. Sawtooth National Forest, Idaho, prescribed fire study: Effects on prickly currant. In: Ribes lacustre. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: https://www.fs.usda.gov/database/feis/plants/shrub/riblac/all.html#FireCaseStudies []. REFERENCE: Lyon, L. Jack. 1971. Vegetal development following prescribed burning of Douglas-fir in south-central Idaho. Res. Pap. INT-105. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 30 p. [42]. SEASON/SEVERITY CLASSIFICATION: Summer/severe STUDY LOCATION: The study site is located in Neal Canyon, 6 miles (9.6 km) north of Ketchum, Idaho, in the Sawtooth National Forest. PREFIRE VEGETATIVE COMMUNITY: The study site was selectively logged in 1950 and 1960.  The remaining trees were primarily pole and sapling Douglas-fir (Pseudotsuga menziesii), many of which were diseased.  Minor tree species included lodgepole pine (Pinus contorta), subalpine fir (Abies lasiocarpa), Engelmann spruce (Picea engelmannii), and quaking aspen (Populus tremuloides).  Understory vegetation was dominated by Rocky Mountain maple (Acer glabrum) and mountain snowberry (Symphoricarpos oreophilus). Prickly currant was a minor species. Vegetation layers over and under 18 inches (46 cm) in height were sampled and described separately.  Prickly currant under 18 inches occurred on 20 percent of the study quadrats.  Prickly currant abundance of plants over 18 inches in height was: density                     0.9 plant/1,000 sq feet canopy cover                0.67 percent aerial crown volume        16.8 cubic feet/1,000 sq feet TARGET SPECIES PHENOLOGICAL STATE: Prickly currant was probably in flowering and fruiting stages on August 1, the day of the fire.  SITE DESCRIPTION: The mountainous study site is at 6,500 feet (1,980 m) elevation.  Annual precipitation is from 14 to 17 inches (360-430 mm), most occurring in the winter as snow.  The slope averages 64 percent and the aspect is 10 to 20 degrees.  The soils are rocky, averaging 50 percent gravel. FIRE DESCRIPTION: The fire was started at 8 a.m. on August 1, 1963 and burned until 5 p.m. Air temperature was 50 degrees Fahrenheit (10 deg C) in the morning and nearly 80 degrees Fahrenheit (27 deg C) in the afternoon.  Relative humidity declined from 50 percent in the morning to 10 percent in the afternoon.  Fuel moisture sticks indicated 5 to 6 percent moisture. Surface winds were less than 5 miles per hour, but fire-induced gusts up to 30 miles per hour were recorded. The fire was severe and crowned.  All litter, herbaceous plants, dead woody stems less than 3 inches (7.6 cm) in diameter, and live woody stems less than 2 inches (5.1) in diameter were consumed.  The Douglas-fir overstory was killed.  The mean heat flux, measured using water-can integrating devices, was 200 calories per second for 45 seconds. FIRE EFFECTS ON TARGET SPECIES: Prickly currant was killed by the fire.  Prickly currant was observed on the study site beginning in postfire year 2.  However, no prickly currant less than 18 inches in height was detected during the 7 postfire years in 2- by 2-foot quadrats.  Density, height, and crown volume for prickly currant over 18 inches in height was originally recorded using quarter-point sampling [79], but this technique was eventually deemed inadequate because of low plant densities.  Beginning in 1966 (postfire year 3), 0.04 acre (0.016 ha) plots were sampled, both in the original quarter-point sampling area and in three additional subplots located at lower, middle, and upper slope sites; measurable quantities of prickly currant occurred only on the lower and upper slope subplots.  Prickly currant over 18 inches in height was first recorded in postfire year 3, presumably growing from seed in the first or second postfire year.  The following data should be interpreted with caution because of very low sample numbers for prickly currant.                                            Postfire year                                                  1      2      3      4      5      6     7 plants/1,000 sq ft   original study area:     -      -      T      T      -     0.3   0.1   subplots (lower slopes): *      *     0.2    0.2    1.0    1.3   1.8   subplots (upper slopes): *      *     0.1    0.5    0.2    0.2   0.5 canopy cover (%)              original study area:     -      -      T     0.01    -    0.05    T aerial crown volume    (cu ft/1,000 sq ft)      original study area:     -      -     0.1    0.1     -     1.1    T      subplots (lower slopes): *      *     0.2    0.4    2.6    7.4   4.4   subplots (upper slopes): *      *     0.7    2.0    1.4    1.8   2.5 *Data were not collected Height and crown volume were measured and averaged for the first four prickly currants on the burn to reach 18 inches in height.                                            Postfire year                            1      2      3      4      5      6     7 average height (ft)        -      -     1.7    1.6    1.5    2.4   1.9 Average volume (cu ft)     -      -     2.6    3.1    4.4   11.5   6.2 A wildfire site which burned in 1950 was located near the Neal Canyon study site.  Twenty years after the wildfire, prickly currant averaged 2.6 plants per 1,000 square feet, 0.6 percent cover , and 12.2 cubic feet aerial crown volume per 1,000 square feet.  Based on the Neal Canyon prescribed fire and the wildfire, Lyon [42] projects that prickly currant will reach prefire crown volumes 30 to 40 years after fire. FIRE MANAGEMENT IMPLICATIONS: NO-ENTRY

REFERENCES

SPECIES: Ribes lacustre
REFERENCES:  1.  Allen, Robert B.; Peet, Robert K.; Baker, William L. 1991. Gradient        analysis of latitudinal variation in Southern Rocky Mountain forests.        Journal of Biogeography. 18(2): 123-138.  [14875]  2.  Antos, Joseph A.; Shearer, Raymond C. 1980. Vegetation development on        disturbed grand fir sites, Swan Valley, northwestern Montana. Res. Pap.        INT-251.  Ogden, UT: U.S. Department of Agriculture, Forest Service,        Intermountain Forest and Range Experiment Station. 26 p.  [7269]  3.  Antos, Joseph A.; Zobel, Donald B. 1985. Plant form, developmental        plasticity and survival following burial by volcanic tephra. Canadian        Journal of Botany. 63: 2083-2090.  [12553]  4.  Antos, Joseph A.; Zobel, Donald B. 1986. Habitat relationships of        Chamaecyparis nootkatensis in southern Washington, Oregon, and        California. Canadian Journal of Botany. 64: 1898-1909.  [19168]  5.  Bamberg, Samuel A.; Major, Jack. 1968. Ecology of the vegetation and        soils associated with calcareous parent materials in three alpine        regions of Montana. Ecological Monographs. 38(2): 127-167.  [12554]  6.  Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals,        reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's        associations for the eleven western states. Tech. Note 301. Denver, CO:        U.S. Department of the Interior, Bureau of Land Management. 169 p.        [434]  7.  Bradley, Anne F.; Noste, Nonan V.; Fischer, William C. 1992. Fire        ecology of forests and woodlands of Utah. Gen. Tech. Rep. INT-287.        Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain        Research Station. 128 p.  [18212]  8.  Braun, E. Lucy. 1961. The woody plants of Ohio. Columbus, OH: Ohio State        University Press. 362 p.  [12914]  9.  Clary, Warren P. 1983. Overstory-understory relationships: spruce-fir        forests. In: Bartlett, E. T.; Betters, David R., eds.        Overstory-understory relationships in Western forests. Western Regional        Research Publication No. 1. Fort Collins, CO: Colorado State University        Experiment Station: 9-12.  [3310] 10.  Cormack, R. G. H. 1953. A survey of coniferous forest succession in the        eastern Rockies. Forestry Chronicle. 29: 218-232.  [16458] 11.  Cowan, Ian McTaggart. 1945. The ecological relationships of the food of        the Columbian black-tailed deer, Odocoileus hemionus columbianus        (Richardson), in the c. forest region southern Vancouver Island, British        Columbia. Ecological Monographs. 15(2): 110-139.  [16006] 12.  Crane, M. F.; Fischer, William C. 1986. Fire ecology of the forest        habitat types of central Idaho. Gen. Tech. Rep. INT-218. Ogden, UT: U.S.        Department of Agriculture, Forest Service, Intermountain Research        Station. 85 p.  [5297] 13.  Crouch, Glenn L. 1985. Effects of clearcutting a subalpine forest in        central Colorado on wildlife habitat. Res. Pap. RM-258. Fort Collins,        CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain        Forest and Range Experiment Station. 12 p.  [8225] 14.  Daubenmire, Rexford F.; Daubenmire, Jean B. 1968. Forest vegetation of        eastern Washington and northern Idaho. Technical Bulletin 60. Pullman,        WA: Washington State University, Agricultural Experiment Station. 104 p.        [749] 15.  De Grandpre, Louis; Gagnon, Daniel; Bergeron, Yves. 1993. Changes in the        understory of Canadian southern boreal forest after fire. Journal of        Vegetation Science. 4: 803-810.  [23019] 16.  Despain, Don G. 1973. Vegetation of the Big Horn Mountains, Wyoming, in        relation to substrate and climate. Ecological Monographs. 43(3):        329-355.  [789] 17.  Dittberner, Phillip L.; Olson, Michael R. 1983. 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